Publicações/Temas

Por cronologia | Por temas

Publicações a partir do ano 2000

Taxonomia & História Natural

Siewert, R. R., A. H. B. Rosa, G. Lamas & A. V. L. Freitas. 2022. Lost and Found: Taxonomic Notes on Some Butterflies (Lepidoptera: Papilionoidea) from J.F. Zikán found on F. Dissmann’s Collection. Neotropical Entomology (Online version). [PDF]

Rosa, A. H. B., N. Seraphim, P. A. Machado, P. E. Gueratto, T. Sobral-Souza & A. V. L. Freitas. 2022. Eucorna sanarita (Schaus) (Riodinidae: Riodininae): Distribution, systematic position, and conservation of a threatened Brazilian butterfly in the Atlantic Forest biodiversity hotspot. Journal of Insect Conservation (online). [PDF]

Mota, L. L., A. H. B. Rosa, L. R. Vasconcellos, K. R. Willmott & A. V. L. Freitas. 2022. A new subspecies of Mechanitis lysimnia from southern Amazonia (Nymphalidae: Danainae: Ithomiini). Tropical Lepidoptera Research 32(1): 47-51. [PDF]

Freitas, A. V. L. 2022. Immature stages of Splendeuptychia ambra (Nymphalidae: Euptychiina) and the diversity of immature morphology within Splendeuptychia. Tropical Lepidoptera Research 32(1): 52-58. [PDF]

Nakahara, S., M. Rodríguez-Melgarejo, K. Kleckner, T. Corahua-Espinoza, R. Tejeira, M. Espeland, M. M. Casagrande, E. P. Barbosa, J. See, G. Gallice, G. Lamas & K. R. Willmott. 2022. Systematic Revision of a New Butterfly Genus, Cisandina Nakahara & Espeland, n. gen., with Descriptions of Three New Taxa (Lepidoptera: Nymphalidae: Satyrinae). Insect Systematics and Diversity 6(1): 2; 1–30. [PDF]

Barbosa, E. P., N. Seraphim, G. Valencia, A. M. L. Azeredo-Espin & A. V. L. Freitas. 2022. Phylogenetic systematics of Yphthimoides Forster, 1964 and related taxa, with notes on the biogeographical history of Yphthimoides species. Molecular Phylogenetics and Evolution 168: 107390. [PDF]

Freitas, A. V. L., A. H. B. Rosa, K. S. Brown Jr., C. E. B. Nobre, D. H. A. Melo & S. M. Kerpel. 2021. A new and rare Ithomiini from Northeastern Brazil (Lepidoptera: Nymphalidae: Danainae). Journal of the Lepidopterists’ Society 75(1): 44–48. [PDF]

A. V. L. Freitas, E. P. Barbosa & J. Y. O. Carreira. 2021. Immature stages and natural history of Yphthimoides borasta (Nymphalidae: Euptychiina). Tropical Lepidoptera Research 31(1): 42-47. [PDF]

Rosa, A. H. B., L. A. R. Leite, R. P. R. Costa & A. V. L. Freitas. 2021. A New Species of Dynamine Hübner, [1919] (Lepidoptera: Nymphalidae: Biblidinae) from the Atlantic Forest in Southeastern Brazil. Neotropical Entomology 50: 948–957. [PDF]

Magaldi, L. M., L. A. Kaminski, N. Seraphim, A. M. Azeredo-Espin, K. L. Silva-Brandão & A. V. L. Freitas. 2021. Hidden in the wing dots: Disentangling mimetic sister species of butterflies (Riodinidae: Stalachtis) with an integrative approach. Zoologischer Anzeiger 294: 92-99. [PDF]

Zacca, T., M. M. Casagrande, O. H. H. Mielke, B. Huertas, A. V. L. Freitas, M. A. Marín, M. Espeland & K. R. Willmott. 2021. A new euptychiine butterfly species from south Brazil and taxonomic rearrangements for Taydebis Freitas, 2013 and Hermeuptychia Forster, 1964 (Lepidoptera: Nymphalidae: Satyrinae). Zootaxa 5023(4): 555-570. [PDF]

Rosa, A. H. B., B. Huertas, K. R. Willmott, E. P. Barbosa, P. A. Machado, O. H. H. Mielke, C. H. P. Canaan & A. V. L. Freitas. 2021. Fifty years without a name: a new species of Splendeuptychia Forster (Lepidoptera: Nymphalidae: Satyrinae). Zootaxa 5061(1): 95-114. [PDF]

Nakahara, S., E. P. Barbosa, I. Nakamura, G. Lamas & A. V. L. Freitas. 2021. Description of a new species of Yphthimoides Forster, 1964 from Peru (Lepidoptera: Nymphalidae: Satyrinae). Tropical Lepidoptera Research 31(3):138-144. [PDF]

Gueratto, P. E., P. A. Machado, T. M. C. Aguiar, E. P. Barbosa, F. M. S. Dias, J. F. Oliveira-Neto, M. M. Casagrande & A. V. L. Freitas. 2020. Identifying Memphis: A comprehensive and comparative description of the immature stages and natural history of Memphis acidalia victoria (H. Druce, 1877) (Lepidoptera: Nymphalidae). Austral Entomology 59(1): 127-141. [PDF]

Freitas, A. V. L. 2020. A new clearwing butterfly from northeastern Brazil (Nymphalidae: Danainae: Ithomiini). Tropical Lepidoptera Research 30(1): 28-32. [PDF]

Mota, L. L., L. A. Kaminski & A. V. L. Freitas. 2020. The tortoise caterpillar: carnivory and armoured larval morphology of the metalmark butterfly Pachythone xanthe (Lepidoptera: Riodinidae). Journal of Natural History 54(5-6): 309-319. [PDF]

Zacca, T., M. M. Casagrande, O. H. H. Mielke, B. Huertas, E. P. Barbosa, A. V. L. Freitas, G. Lamas, M. Espeland, C. Brévignon, S. Nakahara, M. F. Checa & K. R. Willmott. 2020. Systematics of the Neotropical butterfly genus Paryphthimoides (Lepidoptera: Nymphalidae: Satyrinae), with descriptions of seven new taxa. Insects Systematics & Evolution 1: 1-55. [PDF]

Barbosa, E. P., T. M. C. Aguiar, A. H. B. Rosa, T. Zacca & A. V. L. Freitas. 2020. A new species of butterfly (Lepidoptera: Nymphalidae) from the highlands of Southeastern Brazil. Zootaxa 4819(1): zootaxa.4819.1.6. [PDF]

Zacca, T., M. M. Casagrande, O. H. H. Mielke, B. Huertas, E. P. Barbosa, A. V. L. Freitas & K. R. Willmott. 2020. Description of Emeryus Zacca, Mielke & Casagrande gen. nov. (Lepidoptera: Nymphalidae) to accommodate three species formerly placed in Paryphthimoides Forster, 1964. Austral Entomology 59(3): 505-523. [PDF]

Zacca, T., M. M. Casagrande, O. H. H. Mielke, B. Huertas, M. Espeland, A. V. L. Freitas, K. R. Willmott, S. Nakahara & G. Lamas. 2020. Revalidation of Vareuptychia Forster, 1964, description of Vanima gen. nov., and notes on Euptychia cleophes Godman & Salvin, 1889 (Lepidoptera: Nymphalidae: Satyrinae). Zootaxa 4858(1): 1-34. [PDF]

Freitas, A. V. L., R. B. Francini, O. H. H. Mielke, A. H. B. Rosa, L. M. Magaldi & K. L. Silva-Brandão. 2020. A New and Rare Actinote Hübner (Lepidoptera: Nymphalidae: Heliconiinae: Acraeini) from Southeastern Brazil. Neotropical Entomology 49: 696-703. [PDF]

Freitas, A. V. L., A. H. B. Rosa, C. E. B. Nobre, D. H. A. Melo, L. L. Mota, K. L. Silva-Brandão, P. A. Machado & J. Y. O. Carreira. 2020. Immature Stages, Natural History, Systematics and Conservation of an Endangered Neotropical Butterfly: the Case of Scada karschina delicata (Nymphalidae: Ithomiini). Neotropical Entomology 49(5): 685-695. [PDF]

Freitas, A. V. L. 2020. Immature stages of the Neotropical skipper Saliana longirostris (Sepp, [1840]) (Lepidoptera: Hesperiidae). Tropical Lepidoptera Research 30(2): 81-85. [PDF]

Dias, F. M. S., R. R. Siewert, A. V. L. Freitas, G. Lamas, L. M. Magaldi, O. H. H. Mielke & M. M. Casagrande. 2019. An integrative approach elucidates the systematics of Sea Hayward and Cybdelis Boisduval (Lepidoptera: Nymphalidae: Biblidinae). Systematic Entomology 44(1): 226 – 250.

Willmott, K. R., M. A. Marín, S. Nakahara, T. Pomerantz, G. Lamas, B. Huertas, M. Espeland, L. Xiao., J. P. W. Hall, J. I. R. Willmott & A. V. L. Freitas. 2019. A revision of the new Andean butterfly genus Optimandes Marín, Nakahara & Willmott, n. gen., with the description of a new species (Nymphalidae: Satyrinae: Euptychiina). Tropical Lepidoptera Research 29(1): 29-44.

Carvalho, M. R. M., E. P. Barbosa & A. V. L. Freitas. 2019. Immature stages of Mechanitis polymnia casabranca (Nymphalidae, Danainae). Tropical Lepidoptera Research 29(1): 12-18.

Freitas, A. V. L., M. S. Woodbury, E. P. Barbosa & K. R. Willmott. 2019. A new subspecies of Adelpha messana (C. Felder & R. Felder) from the Brazilian Atlantic forest (Nymphalidae: Limenitidinae). Tropical Lepidoptera Research 29(2): 62-66.

Freitas, A. V. L., R. R. Ramos, K. L. Silva-Brandão, N. Coutoné, L. M. Magaldi, J. L. Pablos, N. Rosser & K. S. Brown Jr. 2019. A New Subspecies of Heliconius hermathena (Nymphalidae: Heliconiinae) from Southern Amazonia. Neotropical Entomology 48(3): 467-475.

Freitas, A. V. L., L. L. Mota, E. P. Barbosa & J. Y. O. Carreira. 2019. Immature stages of the Selaginella-feeding Euptychia mollina (Nymphalidae: Satyrinae) from the Brazilian Amazon. Zoologia, 36: e32053. [PDF]

Freitas, A. V. L., L. L. Mota, T. Zacca & E. P. Barbosa. 2019. Description of a new and highly distinctive genus and species of Euptychiina (Lepidoptera: Nymphalidae: Satyrinae) from the Brazilian southern Amazon. Revista Brasileira de Entomologia 63(3): 254-261.

Freitas, A. V. L., P. E. Gueratto & A. R. Nascimento. 2019. New records of high altitude Actinote (Nymphalidae: Heliconiinae) from central Brazil extend the known distribution by 650 km. Journal of the Lepidopterists’ Society 73(3): 208-210.

Nakahara, S., P. Matos-Maraví, E. P. Barbosa, K. R. Willmott, G. Lamas & A. V. L. Freitas. 2019. Two new species of Taygetina with a possible case of ‘juxta loss’ in Butterflies (Lepidoptera: Nymphalidae: Satyrinae). Insect Systematics and Diversity 3(6): 1-13.

Rosa, A. H. B., P. A. Machado, K. L. Silva-Brandão, M. M. De Souza, P. P. Gomes, M. P. Martins & A. V. L. Freitas. 2019. New geographical records for the critically endangered butterfly Parides panthonus castilhoi (Papilionidae: Troidini). Journal of the Lepidopterists’ Society 73(4): 279-281.

Nakahara, S., G. Lamas, S. Tyler, M. A. Marín, B. Huertas, K. R. Willmott, O. H. H. Mielke & M. Espeland. 2019. A revision of the new genus Amiga Nakahara, Willmott & Espeland, gen. n., described for Papilio arnaca Fabricius, 1776 (Lepidoptera, Nymphalidae, Satyrinae). ZooKeys 821: 85–152. [PDF]

Garraffoni, A. R. S., A. Kieneke, M. Kolicka, P. H. C. Cargosinho, J. Prado, S. S. Nihei & A. V. L. Freitas. 2019. ICZN Declaration 45: a remedy for the nomenclatural and typification dilemma regarding soft-bodied meiofaunal organisms? Marine Biodiversity 49: 2199–2207. [PDF]

Braga, L., Ramos, R. R.; Aguiar, T. M. C. & Freitas, A. V. L. 2018. Distribution Extension of Godartiana byses (Nymphalidae: Satyrinae) and First Record for the State of São Paulo, Brazil. The Lepidopterists’ Society 72(2): 176-179. [PDF]

Freitas A.V.L., Barbosa E.P., Zacca T., Marín M.A., Beirão M.V., Silva A.R.M., Casagrande M., Espeland M. & Willmott K.R. 2018. Before it is too late: description of a new genus and species of butterfly from a highly threatened Brazilian biome. Revista Brasileira de Entomologia 62: 148–158. [PDF]

Freitas A.V.L., Francini R.B., Paluch M., Barbosa E.P. 2018. A new species of Actinote Hübner (Nymphalidae: Heliconiinae: Acraeini) from southeast Brazil. Revista Brasileira de Entomologia 62: 135-147.

Benmesbah M., Zacca T., Casagrande M.M., Mielke O.H.H., Lamas G. & Willmott K.R. 2018. Taxonomic notes on Papilio ocypete Fabricius, 1776 and Papilio helle Cramer, 1779 with description of two new similar species from South America (Lepidoptera: Nymphalidae: Satyrinae). Zootaxa 4425: 115-145. [PDF]

Barbosa E.P., Siewert R.R., Mielke O.H.H., Lamas G., Willmott K.R. & Freitas A.V.L. 2018. Redescription of Yphthimoides patricia (Hayward, 1957), with taxonomic notes on the names Euptychia saltuensis Hayward, 1962 and Yphthimoides manasses (C. Felder & R. Felder, 1867) (Nymphalidae: Satyrinae). – Zootaxa 4422(4): 537-557. [PDF]

Pyrcz T. W., Freitas A. V. L., Boyer P., Dias F. M. S., Dolibaina D. R., Barbosa E. P., Magaldi L. M., Mielke O. H. H., Casagrande M. M. & Lorenc-Brudecka J. 2018. Uncovered Diversity of a Predominantly Andean Butterfly Clade in the Brazilian Atlantic Forest: a Revision of the Genus Praepedaliodes Forster (Lepidoptera: Nymphalidae, Satyrinae, Satyrini). Neotropical Entomology 47: 211–255. [PDF]

Freitas, A. V. L., A. H. B. Rosa, J. Y. O. Carreira, P. E. Gueratto, J. P. Santos, A. Tacioli. 2018. Immature stages of two Moneuptychia from southeastern Brazil (Nymphalidae: Euptychiina). Tropical Lepidoptera Research 28(2): 100-105. [PDF]

Freitas, A. V. L. 2018. Immature stages of the Neotropical skipper Lychnuchoides ozias ozias (Hewitson, 1878) (Lepidoptera: Hesperiidae). Tropical Lepidoptera Research 28: 25-28. [PDF]

Zacca. T., M. M. Casagrande, O. H. H. Mielke, B. Huertas, E. P. Barbosa, A. V. L. Freitas, L. M. Magaldi, M. Espeland, S. Nakahara & K. R. Willmott. 2018. Systematics of the butterfly genus Cissia Doubleday, 1848 (Lepidoptera: Nymphalidae: Satyrinae) using an integrative approach. Arthropod Systematics & Phylogeny 76: 349-376.

Bächtold, A., L. A. Kaminski, L. M. Magaldi, P. S. Oliveira, K. Del-Claro, D. H. Janzen, J. M. Burns, N. Grishin, M. Hajibabaei, W. Hallwachs & A. V. L. Freitas. 2017. Integrative data helps the assessment of a butterfly within the Udranomia kikkawai species complex (Lepidoptera: Hesperiidae): Immature stages, natural history, and molecular evidence. Zoologischer Anzeiger 266: 169–176.

Garraffoni, A. R. S. & A. V. L. Freitas. 2017. Photos belong in the taxonomic Code. Science 355(6327): 805.

Shirai, L. T., E. P. Barbosa, R. R. Greve, L. M. Magaldi, A. R. Nascimento & A. V. L. Freitas. 2017. Natural History of Selenophanes cassiope guarany (Lepidoptera: Nymphalidae: Brassolini): an integrative approach, from molecules to ecology. Annals of the Entomological Society of America 110(2): 145-159.

Zacca, T., M. Paluch, R. Siewert, A. V. L. Freitas, E. P. Barbosa, O. H. H. Mielke & M. M. Casagrande. 2017. Revision of Godartiana Forster (Lepidoptera: Nymphalidae), with the description of a new species from northeastern Brazil. Austral Entomology 56(2): 169-190.

Nakahara, S., E. P. Barbosa & A. V. L. Freitas. 2017. A potentially endangered new species of Euptychia Hübner, 1818 (Lepidoptera: Nymphalidae: Satyrinae) from the Atlantic Coastal Forest of Brazil. Neotropical Entomology 46(3): 302-309.

Moraes, S. S., L. S. Otero & A. V. L. Freitas. 2017. Natural history and comparative morphology of immatures of Gamelia anableps (C. Felder & R. Felder) (Lepidoptera: Saturniidae, Hemileucinae). Neotropical Entomolgy 46(4): 397-408.

Zacca, T., M. Paluch, R. Siewert, A. V. L. Freitas, E. P. Barbosa, O. H. H. Mielke & M. M. Casagrande. 2017. Corrigendum. Austral Entomology 56(3): 349.

Kaminski, L. A., C. J. Callaghan, N. Seraphim, L. M. Magaldi, L. Volkmann & A. V. L. Freitas. 2017. Sertania gen. nov., a new genus of butterflies (Lepidoptera: Riodinidae) from the South American dry diagonal. Zootaxa 4312 (1): 165-179.

Freitas, A. V. L. 2017. Immature stages of the Neotropical satyrine butterfly Taygetis acuta (Nymphalidae: Euptychiina). Tropical Lepidoptera Research 27(1): 1-5.

Barbosa, E. P., M. Marín, C. E. Giraldo, S. Uribe & A. V. L. Freitas. 2016. Description of two new species of the Neotropical genus Yphthimoides Forster, 1964 (Lepidoptera: Nymphalidae: Satyrinae) from the ‘renata clade’. Neotropical Biodiversity 2(1): 87-98.

Freitas, A. V. L., E. P. Barbosa, K. R. Willmott, N. Wahlberg & G. Lamas. 2016. ‘Species’ from two different butterfly genera combined into one: description of a new genus of Euptychiina (Nymphalidae: Satyrinae) with unusually variable wing pattern. Revista Brasileira de Entomologia 60 (2): 157-165.

Freitas, A. V. L., J. Y. O. Carreira, J. P. Santos & E. P. Barbosa. 2016. Immature stages and natural history of two species of Forsterinaria from southeastern Brazil (Lepidoptera: Nymphalidae). Tropical Lepidoptera Research 26(1): 13-18. [PDF]

Freitas, A. V. L., E. P. Barbosa & M. A. Marín. 2016. Immature stages and natural history of the neotropical satyrine Pareuptychia ocirrhoe interjecta (Nymphalidae: Euptychiina). Journal of the Lepidopterists’ Society 70(4): 271-276. [PDF]

Freitas, A. V. L., L. M. Magaldi & K. R. Willmott. 2016. A new subspecies of Oleria gunilla (Nymphalidae: Danainae) from north Mato Grosso, Brazil. Journal of the Lepidopterists’ Society 70(4): 295-301.

Kaminski, L. A., C. A. Iserhard & A. V. L. Freitas. 2016. Thisbe silvestre sp. nov. (Lepidoptera: Riodinidae): a new myrmecophilous butterfly from the Brazilian Atlantic Forest. Austral Entomology, 55(2): 138-146.

Mota, L. L., A. K. Silva, A. V. L. Freitas & L. A. Kaminski. 2016. Immature stages of Archonias brassolis tereas (Godart) (Pieridae: Pierini), with notes on interspecific interactions between mistletoe butterflies. Journal of the Lepidopterists’ Society 70(4): 289-294.

Nakahara, S., E. P. Barbosa, M. A. Marín, A. V. L. Freitas, T. Pomerantz & K. R. Willmott. 2016. Graphita gen. nov., a new genus for Neonympha griphe C. Felder & R. Felder, 1867 (Lepidoptera, Nymphalidae, Satyrinae). Neotropical Entomology 45(6): 675-691.

Willmott, K. R., O. H. H. Mielke & A. V. L. Freitas. 2016. A new subspecies and validation of the enigmatic species Adelpha herbita Weymer endemic to the Brazilian Atlantic forest (Nymphalidae: Limenitidinae). Tropical Lepidoptera Research 26(1): 32-37.

Barbosa, E. P., A. K. Silva, M. Paluch, A. M. L. Azeredo-Espin & A. V. L. Freitas. 2015. Uncovering the hidden diversity of the Neotropical butterfly genus Yphthimoides Forster (Nymphalidae: Satyrinae): description of three new species based on morphological and molecular data. Organisms Diversity and Evolution, 15(3): 577-589.

Freitas, A. V.L. 2015. The taxonomic identity of Heliconius melpomene f. pyritosa var. fumigata Zikán (Lepidoptera: Nymphalidae). Revista Brasileira de Entomologia 59(2): 124–125.

Freitas, A. V.L., T. M. Lewinsohn, L. M. Magaldi & N. Wahlberg. 2015. A new subspecies of Anthanassa (Nymphalidae: Nymphalinae: Melitaeini) from southeastern Brazil. Journal of the Lepidopterists’ Society 69(2): 83-90.

Freitas, A. V. L., E. P. Barbosa, R. R. Siewert, O. H. H. Mielke, T. Zacca & A. M. L. Azeredo-Espin. 2015. Four new species of Moneuptychia (Lepidoptera: Satyrinae: Euptychiina) from Brazil. Zootaxa, 3981 (4): 521–541.

Kaminski, L.A.; Ramos, G.R.; Seraphim, N.; Wahlberg, N.; Marini-Filho, O.J.; Freitas, A.V.L. 2015. Natural history and systematic position of Rhetus belphegor (n.comb) (Lepidoptera:Riodinidae), an endangered butterfly with narrow distribution in Southeast Brazil. Journal of Insect Conservation, 19: 1141-1151.

Cosmo L. G., Barbosa E. P. & Freitas A. V. L. 2014. Biology and morphology of the immature stages of Hermeuptychia atalanta (Lepidoptera: Nymphalidae: Satyrinae). Annales de la Société entomologique de France (N.S.) 50 (1): 82-88

Freitas, A. V. L. & L. A. Kaminski. 2014. Ovipositing off the host plant by two tropical nymphalid butterflies. Journal of the Lepidopterists’ Society 68(2): 143-144.

Freitas, A. V. L., L. A. Kaminski, C. A. Iserhard, L. M. Magaldi, N. Wahlberg, K. L. Silva-Brandão & O. H. H. Mielke. 2014. Paulogramma hydarnis (n. comb.) (Nymphalidae: Biblidinae): distribution, systematic position, and conservation status of a rare and endangered butterfly. Neotropical Entomology 43 (3): 218-226.

Kaminski, L. A., L. L. Mota & A. V. L. Freitas. 2014. Larval cryptic coloration and mistletoe use in the metalmark butterfly Dachetola azora (Lepidoptera: Riodinidae). Entomologica Americana 120(1): 18–23.

Leite, L. A. R., A. V. L. Freitas, E. P. Barbosa, M. M. Casagrande & O. H. H. Mielke. 2014. Immature stages of nine species of genus Dynamine Hübner, [1819]: morphology and natural history (Lepidoptera: Nymphalidae, Biblidinae). SHILAP Revista de Lepidopterologia 42(165): 27-55.

Mota, L. L., L. A. Kaminski & A. V. L. Freitas. 2014. Last instar larvae and pupae of Ourocnemis archytas and Anteros formosus (Lepidoptera: Riodinidae), with a summary of known host plants for the tribe Helicopini. Zootaxa 3838(4): 435-444.

Freitas, A. V. L., E. P. Barbosa, J. P. Santos & O. H. H. Mielke. 2013. A new genus, Atlanteuptychia gen. nov., for Euptychia ernestina (Lepidoptera: Nymphalidae: Satyrinae. Zoologia 30(6): 661-668.

Giraldo, C. E., E. P. Barbosa & A. V. L. Freitas. 2013. Immature stages of Pagyris cymothoe cymothoe (Hewitson, 1855) (Lepidoptera, Danainae, Ithomiini). Tropical Zoology 26(4): 145-153.

Kaminski, L. A., L. L. Mota, A. V. L. Freitas & G. R. P. Moreira. 2013. Two ways to be a myrmecophilous butterfly: natural history and comparative immature-stage morphology of two species of Theope (Lepidoptera: Riodinidae). Biological Journal of the Linnean Society, 108: 844-870.

Siewert, R. R., T. Zacca, F. M. S. Dias, A. V. L. Freitas, O. H. H. Mielke & M. M. Casagrande. 2013. The “Taygetis ypthima species group” (Lepidoptera, Nymphalidae, Satyrinae): taxonomy, variation and description of a new species. Zookeys 356: 11-29.

Zacca, T., O. H. H. Mielke, T. W. Pyrcz, M. M. Casagrande, A. V. L. Freitas & P. Boyer. 2013. Stegosatyrus, a new genus of Euptychiina from the grasslands of neotropical realm (Lepidoptera: Nymphalidae: Satyrinae). Zootaxa 3682 (2): 331–350.

Freitas, A. V. L., L. A. Kaminski, O. H. H. Mielke, E. P. Barbosa & K. L. Silva-Brandão. 2012. A new species of Yphthimoides (Lepidoptera: Nymphalidae: Satyrinae) from the southern Atlantic forest region. Zootaxa, 3526: 31-44.

Freitas, A. V. L., N. Wahlberg, P. F. Matos-Maravi, M. A. Marin & O. H. H. Mielke. 2012. Euptychia boulleti (Le Cerf) n. comb. (Lepidoptera: Nymphalidae: Satyrinae), a Rare and Endangered Butterfly from Southeastern Brazil. Neotropical Entomology, 41(4): 461-467.

Kaminski, L. A., D. Rodrigues & A. V. L. Freitas. 2012. Immature stages of Parrhasius polibetes (Lepidoptera: Lycaenidae): host plants, tending ants, natural enemies, and morphology. Journal of Natural History, 46(11-12): 645-667.

Kaminski, L. A., E. P. Barbosa & A. V. L. Freitas. 2012. Immature stages of the neotropical mistletoe butterfly Cunizza hirlanda planasia Fruhstorfer (Pieridae: Anthocharidini). Journal of the Lepidopterists’ Society, 66(3): 143-146.

Kaminski, L. A., L. L. Mota & A. V. L. Freitas. 2012. Life history of Porthecla ravus (Druce) (Lepidoptera: Lycaenidae), with discussion on the use of Annonaceae by Eumaeini butterflies. Annales de la Société Entomologique de France, 48(3-4): 309-312.

Leite, L. A. R., M. M. Casagrande, O. H. H. Mielke & A. V. L. Freitas. 2012. Immature stages of the Neotropical butterfly, Dynamine agacles agacles. Journal of Insect Science, 12, Article 37: 1-12.

Moraes, A. R., H. F. Greeney, P. S. Oliveira, E. P. Barbosa & A. V. L. Freitas. 2012. Morphology and behavior of the early stages of the skipper, Urbanus esmeraldus, on Urera baccifera, an ant–visited host plant. Journal of Insect Science, 12, Article 52: 1-18.

Francini, R. B., E. P. Barbosa & A. V. L. Freitas. 2011. Immature stages of Actinote zikani (Nymphalidae: Heliconiinae), a critically endangered butterfly from southeastern Brazil. Tropical Lepidoptera Research 21(1): 20-26.

Freitas, A. V. L., O. H. H. Mielke, A. Moser, K. L. Silva-Brandão & C. A. Iserhard. 2011. A new genus and species of Euptychiina (Lepidoptera: Nymphalidae, Satyrinae) from south Brazil. Neotropical Entomology, 40(2): 231-237.

Freitas, A. V. L. 2011. Local polyphagy in Theope lycaenina Bates, 1868 (Riodinidae: Nymphidiini). Journal of the Lepidopterists’ Society, 65(4): 264-265.

Silva, P. L., N. P. Oliveira, E. P. Barbosa, Y. Okada, L. A. Kaminski & A. V. L. Freitas. 2011. Immature stages of the Brazilian crescent butterfly Ortilia liriope (Cramer) (Lepidoptera: Nymphalidae). Neotropical Entomology, 40(3): 322-327.

Barbosa, E. P., L. A. Kaminski & A. V. L. Freitas. 2010. Immature stages of the butterfly Diaethria clymena janeira (Lepidoptera: Nymphalidae: Biblidinae). Zoologia. 27(5): 696-702.

Francini, R. B. & A. V. L. Freitas. 2010. Aggregated oviposition in Actinote pellenea pellenea Hübner (Lepidoptera: Nymphalidae). The Journal of Research on the Lepidoptera 42: 74-78.

Freitas, A. V. L., E. O. Emery & O. H. H. Mielke. 2010. A new species of Moneuptychia Forster (Lepidoptera: Satyrinae: Euptychiina) from Central Brazil. Neotropical Entomology 39(1): 83-90.

Freitas, A. V. L., Barbosa, E. P. & R. B. Francini. 2010. Immature stages of the butterfly Actinote conspicua (Nymphalidae: Heliconiinae: Acraeini). Tropical Lepidoptera Research, 20(2): 100-104.

Kaminski, L. A. & A. V. L. Freitas. 2010. Natural history and morphology of immature stages of the butterfly Allosmaitia strophius (Godart) (Lepidoptera: Lycaenidae) on flower buds of Malpighiaceae. Studies on Neotropical Fauna and Environment, 45(1): 11-19.

Kaminski, L. A. & A. V. L. Freitas. 2009. Fabaceae, a new host plant family for Hypanartia and for the Neotropical Nymphalinae (Lepidoptera: Nymphalidae). The Journal of Research on the Lepidoptera, 41: 76-77.

Freitas, A. V. L., L. A. Kaminski, R. G. Matos & K. L. Silva-Brandão. 2009. Immature stages of the Andean butterfly Actinote rufina (Nymphalidae: Heliconiinae: Acraeini). Tropical Lepidoptera Research 19(1): 18-21.

Freitas, A. V. L., R. B. Francini & T. S. Souza. 2009. Immature stages and natural history of the threatened butterfly Actinote quadra (Nymphalidae: Heliconiinae: Acraeini). Tropical Lepidoptera Research 19(2): 82-88.

Willmott, K. R., A. V. L. Freitas, J. P. W. Hall, K. L. Silva-Brandão & M. Paluch. 2009. A new species of Actinote Hübner from the Eastern Andes of Ecuador (Lepidoptera: Nymphalidae: Heliconiinae). Proceedings of the Entomological Society of Washington, 111(1): 47-56.

Algarin, C. M., M. A. Marin, S. Uribe & A. V. L. Freitas. 2008. Tabla de vida de Pseudodirphia pallida (Lepidoptera: Saturniidae) en condiciones de laboratório. Revista Colombiana de Entomología, 34(1): 116-120.

Freitas, A. V. L. & K. S. Brown Jr. 2008. Immature stages of Ithomia salapia ardea (Lepidoptera: Nymphalidae, Ithomiinae). Tropical Lepidoptera Research, 18(1): 17-19.

Freitas, A. V. L. 2008. Description of the early stages of Leucidia (Lepidoptera, Pieridae). Tropical Lepidoptera Research, 18(1): 30-31.

Freitas, A. V. L. & K. S. Brown Jr. 2008. Immature stages of Vila emilia (Nymphalidae: Biblidinae). Tropical Lepidoptera Research, 18(2): 74-77.

Kaminski, L. A. & A. V. L. Freitas. 2008. Immature stages of the butterfly Magneuptychia libye (Linnaeus) (Lepidoptera: Nymphalidae: Satyrinae). Neotropical Entomology, 37(2): 169-172.

Freitas, A. V. L. 2007. A new species of Moneuptychia Forster (Lepidoptera: Satyrinae: Euptychiina) from the highlands of Southeastern Brazil. Neotropical Entomology, 36(6): 919-925.

Freitas, A. V. L. & C. Peña. 2006. Description of Genus Guaianaza for “Euptychiapronophila (Lepidoptera: Nymphalidae: Satyrinae) with a description of the immature stages. Zootaxa, 1163: 49-59.

Freitas, A. V. L. 2006. Immature Stages of Adelpha malea goyama Schaus (Lepidoptera: Nymphalidae: Limenitidinae). Neotropical Entomology, 35(5): 625-628.

Freitas, A. V. L. & K. S. Brown Jr. 2005. Immature stages of Napeogenes sulphurina Bates, 1862 (Lepidoptera, Nymphalidae, Ithomiinae) from Northeastern Brazil. Journal of the Lepidopterists’ Society, 59(1): 35-37.

Francini, R. B., A. V. L. Freitas & C. M. Penz. 2004. Two new species of Actinote (Lepidoptera, Nymphalidae) from Southeastern Brazil. Zootaxa, 719: 1-10.

Freitas, A. V. L. 2004. A new species of Yphthimoides (Nymphalidae, Satyrinae) from southeastern Brazil. Journal of the Lepidopterists’ Society, 58(1): 7-12.

Freitas, A. V. L. 2004. Immature stages of Amphidecta reynoldsi (Nymphalidae: Satyrinae). Journal of the Lepidopterists’ Society, 58(1): 53-55.

Freitas, A. V. L. 2003. Description of a new genus for “Euptychia peculiaris (Nymphalidae: Satyrinae): Immature stages and systematic position. Journal of the Lepidopterists’ Society, 57(2): 100-106.

Freitas, A. V. L. & K. S. Brown Jr. 2002. Immature stages of Sais rosalia (Nymphalidae, Ithomiinae). Journal of the Lepidopterists’ Society, 56(2): 104-106.

Freitas, A. V. L., D. Murray & K. S. Brown Jr. 2002. Immatures, natural history and the systematic position of Bia actorion (Nymphalidae). Journal of the Lepidopterists’ Society, 56(3): 117-122.

Freitas, A. V. L. 2002. Immature stages of Eteona tisiphone (Nymphalidae: Satyrinae). Journal of the Lepidopterists’ Society, 56(4): 286-288.

Freitas, A. V. L., K. S. Brown Jr. & A. Aiello. 2001. Biology of Adelpha mythra feeding on Asteraceae, a novel plant family for the Neotropical Limenitidinae (Nymphalidae), and new data on Adelpha “Species-Group VII”. Journal of the Lepidopterists’ Society, 54(3): 97-100.


Genética, Filogenia & Evolução

Moraes, S. S., P. E. Gueratto, J. P. dos Santos, M. H. Santos, A. V. L. Freitas & M. Duarte. 2022. Niche modelling and comparative morphology untangle taxonomy of the Dysschema eurocilia clade (Lepidoptera: Erebidae) and reveal a relictual Pleistocene Arc distribution. Systematics and Biodiversity 20:1. [PDF]

Barbosa, E. P., N. Seraphim, G. Valencia, A. M. L. Azeredo-Espin & A. V. L. Freitas. 2022. Phylogenetic systematics of Yphthimoides Forster, 1964 and related taxa, with notes on the biogeographical history of Yphthimoides species. Molecular Phylogenetics and Evolution 168: 107390. [PDF]

Marín, M. A., A. López-Rubio, A. Clavijo, T. W. Pyrcz, A. V. L. Freitas, S. I. Uribe & C. F. Álvarez. 2021. Use of species delimitation approaches to tackle the cryptic diversity of an assemblage of high Andean butterflies (Lepidoptera: Papilionoidea). Genome. [PDF]

Pablos, J. L., A. K. Silva, N. Seraphim, L. M. Magaldi, A. P. Souza, A. V. L. Freitas & K. L. Silva-Brandão. 2021. North-south and climate-landscape-associated pattern of population structure for the Atlantic Forest White Morpho butterflies. Molecular Phylogenetics and Evolution 161: 107157. [PDF]

Moraes, S. S., L. Murillo-Ramos, P. A. Machado, H. R. Ghanavi, L. Magaldi, K. L. Silva-Brandão, M. J. Kato, A. V. L. Freitas & N. Wahlberg. 2021. A Double-Edged Sword: Unrecognized cryptic diversity and taxonomic impediment in Eois (Lepidoptera: Geometridae). Zoologica Scripta 2021: 1-14. [PDF]

Moraes, S. S., Y. Montebello, M. A. Stanton, L. F. Yamaguchi, M. J. Kato & A. V. L. Freitas. 2021. Description of three new species of Geometridae (Lepidoptera) using species delimitation in an integrative taxonomy approach for a cryptic species complex. PeerJ 9: e11304. [PDF]

Magaldi, L. M., L. A. Kaminski, N. Seraphim, A. M. Azeredo-Espin, K. L. Silva-Brandão & A. V. L. Freitas. 2021. Hidden in the wing dots: Disentangling mimetic sister species of butterflies (Riodinidae: Stalachtis) with an integrative approach. Zoologischer Anzeiger 294: 92-99. [PDF]

Silva-Brandão, K. L., M. Cirino, L. L. Magaldi, P. E. Gueratto, R. G. Mattos & A. V. L. Freitas. 2021. Subspecies limits and hidden Wolbachia diversity in Actinote pellenea butterflies. Systematics and Biodiversity (online) [PDF]

Chazot, N., [...], A. V. L. Freitas & N. Wahlberg. 2021. Conserved ancestral tropical niche but different continental histories explain the latitudinal diversity gradient in brush-footed butterflies. Nature Communications 12, 5717 [PDF]

Matos-Maraví, P., N. Wahlberg, A. V. L. Freitas, P. DeVries, A. Antonelli & C. M. Penz. 2021. Mesoamerica is a cradle and the Atlantic Forest is a museum of Neotropical butterfly diversity: insights from the evolution and biogeography of Brassolini (Lepidoptera: Nymphalidae). Biological Journal of the Linnean Society 133(3): 704–724. [PDF]

Doré, M., K. R. Willmott, B. Leroy, N. Chazot, J. Mallet, A. V. L. Freitas, J. P. W. Hall, G. Lamas, K. K. Dasmahapatra, C. Fontaine & M. Elias. 2021. Anthropogenic pressures coincide with Neotropical biodiversity hotspots in a flagship butterfly group. Diversity and Distributions 28(12): 2912-2930. [PDF]

Brown, J. L., A. Paz, M. Reginato, C. A. Renata, C. Assis, M. Lyra, M. K. Caddah, J. Aguirre-Santoro, F. d’Horta, F. R. Amaral, R. Goldenberg, K. L. Silva-Brandão, A. V. L. Freitas, M. T. Rodrigues, F. A. Michelangeli, C. Y. Miyaki & A. C. Carnaval. 2020. Seeing the forest through many trees: Multi‐taxon patterns of phylogenetic diversity in the Atlantic Forest hotspot. Diversity and Distributions 26(9): 1160-1176. [PDF]

Massardo, D., N. W. VanKuren, S. Nallu, R. R. Ramos, P. G. Ribeiro, K. L. Silva-Brandão, M. M. Brandão, M. B. Lion, A. V. L. Freitas, M. Z. Cardoso & M. R. Kronforst. The roles of hybridization and habitat fragmentation in the evolution of Brazil’s enigmatic longwing butterflies, Heliconius nattereri and H. hermathena. BMC Biology 18(1): 1-21. [PDF]

Espeland, M., J. W. Breinholt, E. P. Barbosa, M. M. Casagrande, B. Huertas, G. Lamas, M. A. Marín, O. H. H. Mielke, J. Y. Miller, S. Nakahara, D. Tan, A. D. Warren, T. Zacca, A. Y. Kawahara, A. V. L. Freitas & K. R. Willmott. 2019. Four hundred shades of brown: Higher level phylogeny of the problematic Euptychiina (Lepidoptera, Nymphalidae, Satyrinae) based on hybrid enrichment data. Molecular Phylogenetics and Evolution 131: 116 – 124. [PDF]

Chazot, N., K. R. Willmott, G. Lamas, A. V. L. Freitas, F. Piron‐Prunier, C. F. Arias, J. Mallet & D. L. De-Silva. 2019. Renewed diversification following Miocene landscape turnover in a Neotropical butterfly radiation. Global Ecology and Biogeography 28(8): 1118-1132.

Chazot, N., N. Wahlberg, A. V. L. Freitas, C. Mitter, C. Labandeira, J-C. Sohn, R. K. Sahoo, N. Seraphim, R. de Jong &. M. Heikkilä. 2019. Priors and posteriors in Bayesian Timing of Divergence analyses: The age of butterflies revisited. Systematic Biology 68(5): 797-813.

Seraphim, N., Kaminski, L. A., DeVries, P. J., Penz, C., Callaghan, C., Wahlberg, N., Silva-Brandão, K. L., & Freitas, A. V. L. 2018. Molecular phylogeny and higher systematics of the metalmark butterflies (Lepidoptera: Riodinidae). Systematic Entomology. 43: 407-425.

Husch, P.E., Ferreira, D.G., Seraphim, N., Harvey, N., Silva‐Brandão, K.L., Sofia, S.H. & Sosa‐Gómez, D.R. 2018. Structure and genetic variation among populations of Euschistus heros from different geographic regions in Brazil. Entomologia Experimentalis et Applicata 166(3): 191-203.

Silva-Brandão K.L., Peruchi A., Seraphim N., Murad N.F., Carvalho R.A., et al. (2018) Loci under selection and markers associated with host plant and host-related strains shape the genetic structure of Brazilian populations of Spodoptera frugiperda (Lepidoptera, Noctuidae). PLOS ONE 13(5): e0197378. [PDF]

Chazot, N. D. L. De-Silva, K. R. Willmott, A. V. L. Freitas, G. Lamas, J. Mallet, C. E. Giraldo, S. Uribe & M. Elias. 2018. Contrasting patterns of Andean diversification among three diverse clades of Neotropical clearwing butterflies. Ecology and Evolution 8: 3965-3982. [PDF]

Zenker, M. M., N. Wahlberg, G. Brehm, J. A. Teston, L. Przybylowicz, M. R. Pie & A. V. L. Freitas. 2017. Systematics and origin of moths in the subfamily Arctiinae (Lepidoptera, Erebidae) in the Neotropical region. Zoologica Scripta 46(3): 348-362.

De-Silva, D. L., L. L. Mota, N. Chazot, R. Mallarino, K. L. Silva-Brandão, L. M. Gómez-Piñerez, A. V. L. Freitas, G. Lamas, M. Joron, J. Mallet, C. E. Giraldo, S. Uribe, T. Särkinen, S. Knapp, C. D. Jiggins, K. R. Willmott & M. Elias. 2017. North Andean origin and diversification of the largest ithomiine butterfly genus. Scientific Reports 7(45966): 1-17.

Marín, M. A., C. Peña, S. I. Uribe & A. V. L. Freitas. 2017. Morphology agrees with molecular data: phylogenetic affinities of Euptychiina butterflies (Nymphalidae: Satyrinae). Systematic Entomology 42(4): 768-785.

Chazot, N., K. R. Wilmott, A. V. L. Freitas, D. L. De-Silva, T. Pellens & M. Elias. 2016. Patterns of species, phylogenetic and mimicry diversity of clearwing butterflies in the Neotropics. Páginas 333-354. In: Biodiversity Conservation and Phylogenetic Systematics. Pellens, R. & P. Grandcolas (Editors). Springer International Publishing, AG, Switzerland. xvii + 390p.

Chazot, N., K. R. Willmott, F. L. Condamine, D. L. de-Silva, A. V. L. Freitas, G. Lamas, H. Morlon, C. E. Giraldo, C. D. Jiggins, M. Joron, J. Mallet, S. Uribe & M. Elias. 2016. Into the Andes: multiple independent colonizations drive montane diversity in the Neotropical clearwing butterflies Godyridina Molecular Ecology 25 (22): 5765-5784. [PDF]

Seraphim, N., M. A. Barreto, G. S. S. Almeida, A. P. Esperanço, R. F. Monteiro, A. P. Souza, A. V. L. Freitas & K. L. Silva-Brandão. 2016. Genetic diversity of Parides ascanius (Lepidoptera: Papilionidae: Troidini): implications for the conservation of Brazil’s most iconic endangered invertebrate species. Conservation Genetics 17(3): 533-546. [PDF]

Talavera, G., L. A. Kaminski, A. V. L. Freitas & R. Vila. 2016. One-note samba: the biogeographical history of the relict Brazilian butterfly Elkalyce cogina. Journal of Biogeography, 43(4): 727–737.

Garzón-Orduña, I. J., K. L. Silva-Brandão, K. R. Willmott, A. V. L. Freitas & A. V. Z. Brower. 2015. Incompatible ages for clearwing butterflies based on alternative secondary calibrations. Systematic Biology, 65(5): 752-767.

Brower, A. V. Z., K. R. Willmott, K. L. Silva-Brandão, I. J. Garzón-Orduña & A. V. L. Freitas. 2014. Phylogenetic relationships of ithomiine butterflies (Lepidoptera: Nymphalidae: Danainae) as implied by combined morphological and molecular data. Systematics and Biodiversity 12(2): 133-147 (including a “corrigenda”: pp. 259-260).

Seraphim, N., M. A. Marín, A. V. L. Freitas & K. L. Silva-Brandão. 2014. Morphological and molecular marker contributions to disentangling the cryptic Hermeuptychia hermes species complex (Nymphalidae: Satyrinae: Euptychiina). Molecular Ecology Resources 14: 39–49.

Matos-Maravi, P. F., C. Peña, K. R. Willmott, A. V. L. Freitas & N. Wahlberg. 2013. Systematics and evolutionary history of butterflies in the ‘‘Taygetis clade’’ (Nymphalidae: Satyrinae: Euptychiina): Towards a better understanding of Neotropical biogeography. Molecular Phylogenetics and Evolution, 66(1): 54-68.

Penz, C. M., A. V. L. Freitas, L. A. Kaminski, M. M. Casagrande & P. J. DeVries. 2013. Adult and early-stage characters of Brassolini contain conflicting phylogenetic signal (Lepidoptera, Nymphalidae). Systematic Entomology 38: 316-333.

Marín, M. A., C. Peña, A. V. L. Freitas, N. Wahlberg & S. I. Uribe. 2011. From the phylogeny of the Satyrinae butterflies to the systematics of Euptychiina (Lepidoptera: Nymphalidae): History, progress and prospects. Neotropical Entomology 40(1): 1-13.

Peña, C., S. Nylin, A. V. L. Freitas & N. Wahlberg. 2010. Biogeographic history of the butterfly subtribe Euptychiina (Lepidoptera, Nymphalidae, Satyrinae). Zoologica Scripta, 39(3): 243-258.

Elias, M., M. Joron, K. Willmott, K. L. Silva-Brandão, V. Kaiser, C. F. Arias, L. M. Gómez Piñeres, S. Uribe, A.V.Z. Brower, A V. L. Freitas & C. Jiggins. 2009. Out of the Andes: patterns of diversification in clearwing butterflies. Molecular Ecology, 18: 1716-1729. [PDF]

Marín, M. A., A. López, A. V. L. Freitas & S. I. Uribe. 2009. Caracterización molecular de Euptychiina (Lepidoptera: Satyrinae) del norte de la cordillera central de los Andes. Revista Colombiana de Entomología 35(2): 235-244.

Silva-Brandão, K. L., M. L. Lyra & A. V. L. Freitas. 2009. Barcoding Lepidoptera: Current situation and perspectives on the usefulness of a contentious technique. Neotropical Entomology 38(4): 441-451.

Wahlberg, N., J. Leneveu, U. Kodandaramaiah, C. Peña, S. Nylin, A. V. L. Freitas & A. V. Z. Brower. 2009. Nymphalid butterflies diversify following near demise at the Cretaceous/Tertiary boundary. Proceedings of the Royal Society B 276: 4295-4302.

Silva-Brandão, K. L., A. M. L. Azeredo-Espin & A. V. L. Freitas. 2008. New evidence on the systematic and phylogenetic position of Parides burchellanus (Lepidoptera: Papilionidae). Molecular Ecology Resources, 8(3): 502-511.

Silva-Brandão, K. L., N. Wahlberg, R. B. Francini, A. M. L. Azeredo-Espin, K. S. Brown Jr., M. Paluch, D. C. Lees & A. V. L. Freitas. 2008. Phylogenetic relationships of butterflies of the tribe Acraeini (Lepidoptera, Nymphalidae, Heliconiinae) and the evolution of host plant use. Molecular Phylogenetics and Evolution, 46: 515-531.

Wahlberg, N. & A. V. L. Freitas. 2007. Colonization of and radiation in South America by butterflies in the subtribe Phyciodina (Lepidoptera: Nymphalidae). Molecular Phylogenetics and Evolution, 44: 1257-1272.

Brower, A. V. Z., A. V. L. Freitas, M. Lee, K. L. Silva-Brandão, A. Whinnett & K. R. Willmott. 2006. Phylogenetic relationships among the Ithomiini (Lepidoptera: Nymphalidae) inferred from one mitochondrial and two nuclear gene regions. Systematic Entomology 31: 288-301.

Peña, C., N. Wahlberg, E. Weingartner, U. Kodandaramaiah, S. Nylin, A. V. L. Freitas & A. V. Z. Brower. 2006. Higher level phylogeny of Satyrinae butterflies (Lepidoptera: Nymphalidae) based on DNA sequence data. Molecular Phylogenetics and Evolution, 40: 29-49.

Willmott, K. R. & A. V. L. Freitas. 2006. Higher-level phylogeny of the Ithomiinae (Lepidoptera: Nymphalidae): classification, patterns of larval hostplant colonization and diversification. Cladistics, 22(4): 297-368.

Silva-Brandão, K. L., A. V. L. Freitas, A. V. Z. Brower & V. N. Solferini. 2005. Phylogenetic relationships of the New World Troidini swallowtails (Lepidoptera: Papilionidae) based on COI, COII, and EF-1a genes. Molecular Phylogenetics and Evolution 36: 468-483.

Freitas, A. V. L. & K. S. Brown Jr. 2004. Phylogeny of the Nymphalidae (Lepidoptera). Systematic Biology, 53(3): 363-383.


Ecologia, Diversidade & Conservação

Machado, P. A., P. E. Gueratto, S. S. Moraes, A. R. Nascimento, M. J. Kato & A. V. L. Freitas. 2022. Temporal Distribution of Eois Immatures (Lepidoptera, Geometridae) in a Tropical Forest. The Journal of the Lepidopterists’ Society 76(4): 261-267. [PDF]

Rosa, A. H. B., N. Seraphim, P. A. Machado, P. E. Gueratto, T. Sobral-Souza & A. V. L. Freitas. 2022. Eucorna sanarita (Schaus) (Riodinidae: Riodininae): Distribution, systematic position, and conservation of a threatened Brazilian butterfly in the Atlantic Forest biodiversity hotspot. Journal of Insect Conservation (online). [PDF]

Shirai, L. T., M. A. Stanton, G. B. D’Angelo, T. B. Conforti, A. V. L. Freitas, M. J. Kato & L. F. Yamaguchi. 2022. Interaction Gardens and Butterfly Catalogues: a Joint Strategy to Promote Capacity Development in Protected Areas and Reduce the Extinction of Experience in Cities. Cities and the Environment (CATE) 15(1): article 3. [PDF]

Melo, D. H. A., A. V. L. Freitas, M. Tabarelli, B. K. C. Filgueiras & I. R. Leal. 2022. Aridity and chronic anthropogenic disturbance as organizing forces of fruit-feeding butterfly assemblages in a Caatinga dry forest. Biotropica, 00, 1–12. [PDF]

Noll, F. B., M. F. C. Barbosa, E. F. Santos, R. C. Castilho, C. J. E. Lamas, A. V. L. Freitas & G. J. Moraes. 2022. The contribution of the BIOTA/FAPESP Program to the advancement of the knowledge on terrestrial invertebrates. Biota Neotropica 22(spe): e20221398. [PDF]

Lewinsohn, T.M., K. Agostini, A. V. L. Freitas & A. S. Melo. 2022. Insect decline in Brazil: an appraisal of current evidence. Biology Letters 18: 20220219. [PDF]

Freitas, A. V. L., A. H. B. Rosa & B. R. V. Costa. 2021. The watercolors of Brazilian butterflies by Paul-André Robert – a dream eclosing after 60 years. News of The Lepidopterists’ Society 63(4): 163-165. [PDF]

Lourenço, G. M., W. Dáttilo, S. P. Ribeiro & A. V. L. Freitas. 2021. Biological Aspects and Movements of Neotropical Fruit-Feeding Butterflies. Neotropical Entomology 51: 43-53. [PDF]

Ramos, R.R., R. B. Francini, M. E. M. Habib & A. V. L. Freitas. 2021. Seasonal Patterns of Host Plant Use in an Assemblage of Heliconiini Butterflies (Lepidoptera: Nymphalidae) in a Neotropical forest. Neotropical Entomology. [PDF]

Araujo, P. F., A. V. L. Freitas, G. A. S. Gonçalvez & D. B. Ribeiro. 2021. Vertical stratification on a small scale: the distribution of fruit-feeding butterflies in a semi-deciduous Atlantic forest in Brazil. Studies on Neotropical Fauna and Environment 56(1): 10-39. [PDF]

Birskis-Barros, I., A. V. L. Freitas & P. R. Guimarães Jr. 2021. Habitat generalist species constrain the diversity of mimicry rings in heterogeneous habitats. Scientific Reports 11(5072): 1-12. [PDF]

Freitas, A. V. L., D. G. Muniz, J. Y. O. Carreira & P. S. Oliveira. 2021. Fruit-Feeding Butterfly Assemblages in a Neotropical Savanna: Assessing Phenological Patterns Using Baited Traps. The Journal of the Lepidopterists’ Society 75(2): 88-103. [PDF]

Sobral-Souza, T., J. Stropp, J. P. Santos, V. M. Prasniewski, N. Szinwelski, B. Vilela, A. V. L. Freitas, M. C. Ribeiro & J. Hortal. 2021. Knowledge gaps hamper understanding the relationship between fragmentation and biodiversity loss: the case of Atlantic Forest fruit-feeding butterflies. PeerJ 9:e11673. [PDF]

Silva-Brandão, K. L., A. V. L. Freitas, M. Z. Cardoso, R. Cogni & A. B. B. Morais. 2021. The Chemistry and Chemical Ecology of Lepidopterans as Investigated in Brazil. pp. 37-66. In Kinghorn A.D., Falk H., Gibbons S., Asakawa Y., Liu JK., Dirsch V.M. (eds) Progress in the Chemistry of Organic Natural Products, v. 116. Progress in the Chemistry of Organic Natural Products, vol 116. Springer, Cham. [PDF]

Freire, G. B., T. Silva, H. Oliveira, C. Collier, H. P. Rodrigues, J. P. Dias, J. P. Santos, O. J. Marini-Filho, A. V. L. Freitas, A. M. Smilanich, L. A. Dyer & Ivone R. Diniz. 2021. Good Things Come in Larger Packages: Size Matters for Adult Fruit-Feeding Butterfly Dispersal and Larval Diet Breadth. Diversity 13(12): 664. [PDF]

Santos, J. P., T. Sobral-Souza, K. S. Brown Jr., M. H. Vancine, M. C. Ribeiro & A. V. L. Freitas. 2020. Effects of landscape modification on species richness patterns of fruit-feeding butterflies in Brazilian Atlantic Forest. Diversity and Distributions 26(2): 196-208.

Gueratto, P. E., J. Y. O. Carreira, J. P. Santos, A. Tacioli & A. V. L. Freitas. 2020. Effects of forest trails on the community structure of tropical butterflies. Journal of Insect Conservation 24(2): 309-319. [PDF]

Lourenço, G. M., P. Luna, R. Guevara, W. Dattilo, A. V. L. Freitas & S. P. Ribeiro. 2020. Temporal shifts in butterfly diversity: responses to natural and anthropic forest transitions. Journal of Insect Conservation 24(2): 353-363. [PDF]

Rosa, A. H. B., D. B. Ribeiro & A. V. L. Freitas. 2020. Population biology, natural history and conservation of two endangered high elevation Neotropical butterflies. Journal of Insect Conservation 24(4): 681-694. [PDF]

Rosser, N., L. T. Shirai, K. K. Dasmahapatra, J. Mallet & A. V. L. Freitas. 2020. The Amazon river is a suture zone for a polyphyletic group of co‐mimetic heliconiine butterflies. Ecography 43: 1–11. [PDF] [Vídeo]

Nascimento, A.R., I. K. Malinov, G. Freire Jr, A. V. L. Freitas & I. R. Diniz. 2020. The Temporal Dynamics of Two Morpho Fabricius, 1807 Species (Lepidoptera: Nymphalidae) are Affected Differently by Fire in the Brazilian Savanna. Environmental Entomology 49(6): 1449–1454. [PDF]

Filgueiras, B. K. C., D. H. A. Melo, M. Uehara-Prado, A. V. L. Freitas, I. R. Leal & M. Tabarelli. 2019. Compensatory dynamics on the community structure of fruit-feeding butterflies across hyper-fragmented Atlantic forest habitats. Ecological Indicators 98: 276-284.

Iserhard, C. A., L. Duarte, N. Seraphim & A. V. L. Freitas. 2019. How urbanization affects multiple dimensions of biodiversity in tropical butterfly assemblages. Biodiversity and Conservation 28(3): 621-638. [PDF]

Lourenço, G. M., G. R. Soares, T. P. Santos, W. Dattilo, A. V. L. Freitas & S. P. Ribeiro. 2019. Equal but different: Natural ecotones are dissimilar to anthropic edges PLoS ONE 14(3): e0213008. 18pp. [PDF]

Uehara-Prado, M & A. V. L. Freitas. 2019. Population structure of Taygetis ypthima (Nymphalidae, Euptychiina) in Southeast Brazil. Journal of the Lepidopterists’ Society 73(1): 63-64. [PDF]

Rosser, N., A. V. L. Freitas, B. Huertas, M. Joron, G. Lamas, C. Mérot, F. Simpson, K. R. Willmott, J. Mallet & K. K. Dasmahapatra. 2019. Cryptic speciation associated with geographic and ecological divergence in two Amazonian Heliconius butterflies. Zoological Journal of the Linnean Society 186(1): 233-249.

Melo, D. H. A., B. K. C. Filgueiras, C. A. Iserhard, L. Iannuzzi, A. V. L. Freitas & I. R. Leal. 2019. Effect of habitat loss and fragmentation on fruit-feeding butterflies in the Brazilian Atlantic Forest. Canadian Journal of Zoology, 97(7): 588-596. [PDF]

Rosa, A. H. B., L. V. Perillo, F. S. Neves, D. B. Ribeiro & A. V. L. Freitas. 2019. Butterflies collected using malaise traps as useful bycatches for ecology and conservation. Journal of Threatened Taxa 11(9): 14235-14237. [PDF]

Cosmo, L. G.A. R. Nascimento, R. Cogni & A. V. L. Freitas. 2019. Temporal distribution in a tri-trophic system associated with Piper amalago L. in a tropical seasonal forest. Arthropod-Plant Interactions 13(4): 647-652. [PDF]

Waichert, C., T. G. Sobrinho, A. V. L. Freitas, A. H. B. Rosa, C. de A. Sampaio, D. Zeppelini, F. C. Massariol, F. Z. Vaz-de-Melo, F. F. Salles, J. Musiello-Fernandes, K. S. Furieri, K. G. da Costa & V. S. do Amaral. 2019. Invertebrados ameaçados de extinção no estado do Espírito Santo. Páginas 216-229. Capítulo 10. In: Fauna e flora ameaçadas de extinção no estado do Espírito Santo. Fraga, C. N. de, M. de H. Formigoni & F. G. Chaves (organizadores). Instituto Nacional da Mata Atlântica. 432 pp. ISBN 978-65-81414-01-6

de Andrade, R. B., J. K. Balch, J. Y. O. Carreira, P. M. Brando & A. V. L. Freitas. 2017. The impacts of recurrent fires on diversity of fruit-feeding butterflies in a south-eastern Amazon forest. Journal of Tropical Ecology 33 (1): 22-32. [PDF]

Seixas, R. R., S. E. Santos, Y. Okada & A. V. L. Freitas. 2017. Population biology of the sand forest specialist butterfly Heliconius hermathena hermathena (Hewitson) (Nymphalidae: Heliconiinae) in central Amazonia. Journal of the Lepidopterists’ Society 71(3): 133–140.

Santos, J. P., C. A. Iserhard, J. Y. O. Carreira & A. V. L. Freitas. 2017. Monitoring fruit-feeding butterfly assemblages in two vertical strata in seasonal Atlantic Forest: temporal species turnover is lower in the canopy. Journal of Tropical Ecology 33(5): 345-355. [PDF]

Rosa, A. H. B., M. C. Campis, D. B. Ribeiro & A. V. L. Freitas. 2017. New record of the threatened butterfly Drephalys mourei (Hesperiidae) in a heavily disturbed area in Southeastern Brazil. Journal of the Lepidopterists’ Society 71(4): 278-280.

Shirai, L. T., L. L. Mota & A. V. L. Freitas. 2017. Aggregation of Epityches eupompe (Nymphalidae: Ithomiini) in southern Brazil. Tropical Lepidoptera Research 27(2): 111-114.

Freitas, A. V. L. 2017. Biological notes on Utetheisa pulchelloides (Erebidae) in Tuamotu, French Polynesia. Tropical Lepidoptera Research 27(2): 115-116.

Chazot, N., K. R. Wilmott, A. V. L. Freitas, D. L. De-Silva, T. Pellens & M. Elias. 2016. Patterns of species, phylogenetic and mimicry diversity of clearwing butterflies in the Neotropics. Páginas 333-354. In: Biodiversity Conservation and Phylogenetic Systematics. Pellens, R. & P. Grandcolas (Editors). Springer International Publishing, AG, Switzerland. xvii + 390p.

Duarte, L. D. S., V. J. Debastiani, A. V. L. Freitas & V. D. Pillar. 2016. Dissecting phylogenetic fuzzy weighting: theory and application inmetacommunity phylogenetics. Methods in Ecology and Evolution 7(8): 937-946.

Filgueiras, B. K. C., D. H. A. Melo, I. R. Leal, M. Tabarelli, A. V. L. Freitas & Luciana Iannuzzi. 2016. Fruit-feeding butterflies in edge-dominated habitats: community structure, species persistence and cascade effect. Journal of Insect Conservation 20(3): 539-548.

Pinheiro, C. E. G., A. V. L. Freitas, V. C. Campos, P. J, DeVries & C. M. Penz. 2016. Both palatable and unpalatable butterflies use bright colors to signal difficulty of capture to predators. Neotropical Entomology 45(2): 107-113.

Ribeiro, D. B., M. R. Williams, A. Specht & A. V. L. Freitas. 2016. Vertical and temporal variability in the probability of detection of fruit-feeding butterflies and moths (Lepidoptera) in tropical forest. Austral Entomology, 55(1): 112-120.

Santos, J. P., O. J. Marini-Filho, A. V. L. Freitas & M. Uehara-Prado. 2016. Monitoramento de Borboletas: o Papel de um Indicador Biológico na Gestão de Unidades de Conservação. Biodiversidade Brasileira, 6(1): 87-99.

Seraphim, N., M. A. Barreto, G. S. S. Almeida, A. P. Esperanço, R. F. Monteiro, A. P. Souza, A. V. L. Freitas & K. L. Silva-Brandão. 2016. Genetic diversity of Parides ascanius (Lepidoptera: Papilionidae: Troidini): implications for the conservation of Brazil’s most iconic endangered invertebrate species. Conservation Genetics 17(3): 533-546. [PDF]

Silva-Brandão, K. L., D. B. Ribeiro, M. M. Brito, A. J. A. Olavarrieta & A. V. L. Freitas. 2016. The influence of agricultural and forest landscapes on the diversity of Lepidoptera. Páginas 190-205. Capítulo 12. In: Biodiversity in Agricultural Landscapes of Southeastern Brazil. Gheler-Costa, C., M. C. Lyra-Jorge & L. M. Verdade (Editors). De Gruyter Open Ltd, Warsaw/Berlin. XXIV + 342 pp.

Talavera, G., L. A. Kaminski, A. V. L. Freitas & R. Vila. 2016. One-note samba: the biogeographical history of the relict Brazilian butterfly Elkalyce cogina. Journal of Biogeography, 43(4): 727–737.

Zenker, M. M., A. V. L. Freitas, R. Rougerie, M. Laguerre, J. A. Teston & M. R. Pie. 2016. Uncovering tiger moth diversity in Brazil’s most endangered biome. Barcode Bulletin 7(2): 22-23.

Zenker, M. M., R. Rougerie, J. A. Teston, M. Laguerre, M. R. Pie & A. V. L. Freitas. 2016. Fast census of moth diversity in the Neotropics: A comparison of field-assigned morphospecies and DNA barcoding in tiger moths. PLoS ONE 11(2): e0148423.

Kaminski, L. A., R. Dell’Erba, E. P. Barbosa & A. V. L. Freitas. 2015. New distribution records and notes on the habitat of Magneuptychia flavofascia Zacca & Siewert, 2014 (Lepidoptera: Nymphalidae). Checklist, 11(4): article 1692.

Zenker, M. M., P. J. DeVries, C. M. Penz, J. A. Teston, A. V. L. Freitas & M. R. Pie. 2015. Diversity and composition of Arctiinae moth assemblages along elevational and spatial dimensions in Brazilian Atlantic Forest. Journal of Insect Conservation 19(1): 129–140.

Freitas, A. V. L., C. A. Iserhard, J. P. Santos, J. Y. O. Carreira, D. B. Ribeiro, D. H. A. melo, A. H. B. Rosa, O. J. Marini-Filho, G. M. Accacio & M. Uehara-Prado. 2014. Studies with butterfly bait traps: an overview. Revista Colombiana de Entomología 40 (2): 209-218. [PDF]

Gomes, V., G. M. Lourenço, D. Soldati, C. A. Iserhard, T. S. Souza, L. A. Kaminski & A. V. L. Freitas. 2014. New geographical records for the threatened butterfly Actinote quadra (Lepidoptera: Nymphalidae: Heliconiinae). Journal of the Lepidopterists’ Society 68(4): 289-291.

Kerpel, S. M., A. Ferreira-Junior & A. V. L. Freitas. 2014. New record of the endangered Brazilian swallowtail Heraclides himeros baia (Rothschild & Jordan, 1906). Journal of the Lepidopterists’ Society 68(2): 145-146.

Melo, D. H. A., B. K. C. Filgueiras, I. R. Leal & A. V. L. Freitas. 2014. Recent records of Morpho menelaus eberti Fischer (Lepidoptera: Nymphalidae), an endangered butterfly in Northeast Brazil. Tropical Lepidoptera Research 24(2): 102-104.

Pinheiro, C. E. G. & A. V. L. Freitas. 2014. Some possible cases of escape mimicry in Neotropical butterflies. Neotropical Entomology 43(5): 393-398. [PDF]

Ribeiro, D. B. & A. V. L. Freitas. 2014. Brazil’s new laws bug collectors. Science 345(6204): 1571.

Sant’Anna, C. L. B., D. B. Ribeiro, L. C. Garcia & A. V. L. Freitas. 2014. Fruit-feeding butterfly communities are influenced by restoration age in tropical forests. Restoration Ecology 22(4): 480-485.

Greve, R. R., C. Callaghan, L. A. Kaminski & A. V. L. Freitas. 2013. The rediscovery of Joiceya praeclarus Talbot 1928 (Lepidoptera: Riodinidae), more than 80 years after its description. Journal of the Lepidopterists’ Society, 67(1): 56-57.

Herkenhoff, E. V., R. F. Monteiro, A. P. Esperanço & A. V. L. Freitas. 2013. Population biology of the endangered fluminense swallowtail butterfly Parides ascanius (Papilionidae: Papilioninae: Troidini). Journal of the Lepidopterists’ Society, 67(1): 29-34.

Iserhard, C. A., K. S. Brown Jr. & A. V. L. Freitas. 2013. Maximized sampling of butterflies to detect temporal changes in tropical communities. Journal of Insect Conservation, 17: 615-622.

Rodrigues, D. & A. V. L. Freitas. 2013. Contrasting egg and larval performances help explain polyphagy in a florivorous butterfly. Arthropod-Plant Interactions, 7: 159-167.

Almeida, A. C & A. V. L. Freitas. 2012. Lepidoptera. Borboletas e Mariposas do Brasil. Exclusiva Publicações, São Paulo. 208pp. ISBN: 85-61931-05-6 (BOOK)

Bächtold, A., K. Del-Claro, L. A. Kaminski, André V.L. Freitas & P. S. Oliveira. 2012. Natural history of an ant–plant–butterfly interaction in a Neotropical savanna. Journal of Natural History, 46(15-16): 943-954.

Beirão, M. V., F. C. Campos-Neto, I. A. Pimenta & A. V. L. Freitas. 2012. Population biology and natural history of Parides burchellanus (Papilionidae: Papilioninae: Troidini), an endangered Brazilian butterfly. Annals of the entomological Society of America, 105(1): 36-43.

Muniz, D. G., A. V. L. Freitas & P. S. Oliveira. 2012. Phenological relationships of Eunica bechina (Lepidoptera: Nymphalidae) and its host plant, Caryocar brasiliense (Caryocaraceae), in a Neotropical savanna. Studies in Neotropical Fauna and Environment 47(2): 111-118.

Ramos, R. R., D. Rodrigues & A. V. L. Freitas. 2012. Oviposition preference and larval performance in a Heliconius erato phyllis (Lepidoptera: Nymphalidae) population from southeastern Brazil: is there a positive relationship? Journal of Natural History, 46(11-12): 669-681.

Ribeiro, D. B. & A. V. L. Freitas. 2012. The effect of reduced-impact logging on fruit-feeding butterflies in Central Amazon, Brazil. Journal of Insect Conservation, 16: 733-744.

Ribeiro, D. B., R. Batista, P. I. Prado, K. S. Brown Jr. & A. V. L. Freitas. 2012. The importance of small scales to the fruit-feeding butterfly assemblages in a fragmented landscape. Biodiversity and Conservation, 21: 811-827.

Freitas, A. V. L., L. A. Kaminski, C. A. Iserhard, E. P. Barbosa & O. J. Marini-Filho. 2011. The endangered butterfly Charonias theano (Boisduval) (Lepidoptera: Pieridae): current status, threats and its rediscovery in São Paulo state, southeastern Brazil. Neotropical Entomology, 40(6): 669-676.

Freitas, A. V. L. & O. J. Marini-Filho (Organizadores). 2011. Plano de ação nacional para a conservação dos lepidópteros ameaçados de extinção. Série Espécies Ameaçadas nº 13. ICMBio, Brasília, Brasil. 124p. (BOOK) [PDF]

Freitas, A. V. L., A. Camargo, C. A. Iserhard, G. M. Accacio, H. P. Romanowski, J. Galinkin, K. S. Brown Jr, M. Duarte, M. Uehara-Prado, M. Z. Cardoso, M. M. Casagrande, O. H. H. Mielke, O. J. Marini Filho, R. F. Monteiro, R. W. H. Honegger & W. L. Overal. 2011. Parte I. Informações Gerais. Páginas 15-74. In: Plano de ação nacional para a conservação dos lepidópteros ameaçados de extinção. Série Espécies Ameaçadas nº 13. Freitas, A. V. L. & O. J. Marini-Filho (Organizadores). ICMBio, Brasília, Brasil. 124p. (BOOK CHAPTER)

Jorge, L. R., P. Cordeiro-Estrela, L. B. Klaczko, Gilson R. P. Moreira & A. V. L. Freitas. 2011. Host-plant dependent wing phenotypic variation in Heliconius erato. Biological Journal of the Linnean Society, 102: 765-774.

Ribeiro, D. B. & A. V. L. Freitas. 2011. Large-sized insects show stronger seasonality than small-sized ones: a case study of fruit-feeding butterflies. Biological Journal of the Linnean Society, (104): 820–827.

Silva-Brandão, K. L. & A. V. L. Freitas. 2011. Capítulo 8. Interações entre borboletas e plantas. Páginas 171-180. In: Ecologia das Interações Plantas-Animais: Uma abordagem ecológico-evolutiva. Del-Claro, K. & H. M. Torezan-Silingardi (Editores). Technical Books Editora, Rio de Janeiro. 333p. (BOOK CHAPTER)

Freitas, A. V. L. 2010. Impactos potenciais das mudanças propostas no Código Florestal Brasileiro sobre as borboletas. Biota Neotropica, 10(4): 53-57.

Kaminski, L. A., A. V. L. Freitas & P. S. Oliveira. 2010. Interaction between mutualisms: Ant-tended butterflies exploit enemy-free space provided by ant-treehopper associations. The American Naturalist, 176(3): 322-334.

Peña, C., S. Nylin, A. V. L. Freitas & N. Wahlberg. 2010. Biogeographic history of the butterfly subtribe Euptychiina (Lepidoptera, Nymphalidae, Satyrinae). Zoologica Scripta, 39(3): 243-258.

Ribeiro, D. B. & A. V. L. Freitas. 2010. Differences in thermal responses in a fragmented landscape: Temperature affects the sampling of diurnal, but not nocturnal fruit-feeding Lepidoptera. The Journal of Research on the Lepidoptera 42: 1-4.

Ribeiro, D. B., P. I. Prado, K. S. Brown Jr. & A. V. L. Freitas. 2010. Temporal diversity patterns and phenology in fruit-feeding butterflies in the Atlantic Forest. Biotropica, 42(6): 710-716.

Rodrigues, D., L. A. Kaminski, A. V. L. Freitas & P. S. Oliveira. 2010. Trade-offs underlying polyphagy in a facultative ant-tended florivorous butterfly: the role of host plant quality and enemy-free space. Oecologia, 163: 719-728.

Aguiar, A. P., B. F. dos Santos, M. S. Couri, J. A. Rafael, C. Costa, S. Ide, M. Duarte, J. Grazia, C. F. Schwertner, A. V. L. Freitas & C. O. Azevedo. 2009. Insecta. Páginas 131-155. Capítulo 8. In: Estado da Arte e Perspectivas para a Zoologia no Brasil. Rocha, R. M. & W. A. P. Boeger (Editores). Resultados dos Simpósios do XXVII Congresso Brasileiro de Zoologia. Curitiba, Editora UFPR. 296 p. (BOOK CHAPTER)

Kaminski, L. A., S. F. Sendoya, A. V. L. Freitas & P. S. Oliveira. 2009. Ecologia comportamental na interface formiga-planta-herbívoro: interações entre formigas e lepidópteros. Oecologia Brasiliensis, 13(1): 27-44.

Sendoya, S. F., A. V. L. Freitas & P. S. Oliveira. 2009. Egg-laying butterflies distinguish predaceous ants by sight. The American Naturalist 174(1): 134-140.

Tourinho, J. L. & A. V. L. Freitas. 2009. Population biology of Euptoieta hegesia (Nymphalidae: Heliconiinae: Argynnini) in an urban area in Southeastern Brazil. The Journal of Research on the Lepidoptera, 41: 40-44.

Uehara-Prado. M. & A. V. L. Freitas. 2009. The effect of rainforest fragmentation on species diversity and mimicry ring composition of ithomiine butterflies. Insect Conservation and Diversity, 2: 23-28.

Uehara-Prado, M., J. O. Fernandes, A. M. Bello, G. Machado, A. J. Santos, F. Z. Vaz-de-Mello & A. V. L. Freitas. 2009. Selecting terrestrial arthropods as indicators of small-scale disturbance: a first approach in the Brazilian Atlantic Forest. Biological Conservation 142: 1220-1228.

Accacio, G. M., A. V. L. Freitas & M. V. Urso-Guimarães. 2008. Invertebrados. Páginas 99-103 (+ 1 mapa). Capítulo 6.5. In: Diretrizes para a conservação e restauração da biodiversidade no Estado de São Paulo. Rodrigues, R. R., C. A. Joly, M. C. W. Brito, A. Paese, J. P. Metzger, L. Casatti, M. A. Nalon, N. Menezes, N. M. Ivanauskas, V. Bolzani & V. L. R. Bononi (organizadores). Governo do Estado de São Paulo, São Paulo, SP. (BOOK CHAPTER)

Brown, K. S. Jr. & A. V. L. Freitas. 2008. Arawacus aethesa. Página 400; Eresia erysice erysice. Página 412; Heliconius nattereri. Páginas 414-415; Melinaea mnasias thera. Página 418; Napeogenes cyrianassa xanthone. Página 419; Tithorea harmonia caissara. Páginas 428-429; Eurytides iphitas. Página 429; Heraclides himeros baia. Página 430; Heraclides himeros himeros. Página 431; Mimoides lysithous harrisianus. Página 432; Parides ascanius. Página 433; Parides bunichus chamissonia. Página 434; Parides lysander matogrossensis. Páginas 436-437; Parides panthonus castilhoi. Páginas 437-438. In: Livro vermelho da fauna brasileira ameaçada de extinção. Machado, A. B. M., G. M. M. Drummond & A. P. Paglia (Eds.). Brasília, DF: MMA; Belo Horizonte, MG: Fundação Biodiversitas. (BOOK CHAPTER)

Canuto, J. Z., T. A. Nogueira, K. S. Brown Jr. & A. V. L. Freitas. 2008. Lepidoptera, Nymphalidae, Ithomiinae, Scada reckia reckia: suggestion of recent range expansion throughout the State of Espirito Santo, SE Brazil. Checklist, 3(2): 159-160.

Freitas, A. V. L. & K. S. Brown Jr. 2008. Magnastigma julia. Página 401; Actinote quadra. Páginas 402-403; Actinote zikani. Páginas 403-404; Caenoptychia boulletti. Páginas 404-405; Episcada vitrea. Página 411; Grasseia menelaus eberti. Página 413; Hyalyris fiammetta. Página 415; Hyalyris leptalina. Página 416; Hypoleria fallens. Página 417; Paititia neglecta. Página 422; Pessonia epistrophus nikolajewna. Página 424; Polygrapha suprema. Página 425; Scada kasrchina delicata. Página 427; Nirodia belphegor. Páginas 445-446. In: Livro vermelho da fauna brasileira ameaçada de extinção. Machado, A. B. M., G. M. M. Drummond & A. P. Paglia (Eds.). Brasília, DF: MMA; Belo Horizonte, MG: Fundação Biodiversitas. (BOOK CHAPTER)

Ribeiro, D. B., P. I. Prado, K. S. Brown Jr. & A. V. L. Freitas. 2008. Additive partitioning of butterfly diversity in a fragmented landscape: Importance of scale and implications for conservation. Diversity and Distributions, 14: 961-968.

Azevedo, C. O., F. Z. Vaz-de-Mello, M. T. Tavares, A. D. Brescovit, P. de Marco Jr., K. Furieri, K. S. Brown Jr. & A. V. L. Freitas. 2007. Os invertebrados terrestres ameaçados de extinção no estado do Espírito Santo. Páginas 105-119 In: Espécies da fauna ameaçados de extinção no estado do Espírito Santo. Pasamani, M. & S. L. Mendes (organizadores.). IPEMA, Vitória, 140pp. (BOOK CHAPTER)

Brown Jr K. S., A. V. L. Freitas, N. Wahlberg, B. Von Schoultz, A. O. Saura & A. Saura. 2007. Chromosomal evolution in the South American Nymphalidae. Hereditas, 144: 137-148.

Brown Jr K. S., A. V. L. Freitas, B. Von Schoultz, A. O. Saura & A. Saura. 2007. Chromosomal evolution of South American frugivorous butterflies in the Satyroid clade (Nymphalidae: Charaxinae, Morphinae and Satyrinae). Biological Journal of the Linnean Society, 92: 467-481.

Uehara-Prado, M. K. S. Brown Jr. & A. V. L. Freitas. 2007. Species richness, composition and abundance of fruit-feeding butterflies in the Brazilian Atlantic Forest: comparison between a fragmented and a continuous landscape. Global Ecology and Biogeography, 16: 43-54.

Freitas, A. V. L., I. R. Leal, M. Uehara-Prado & L. Iannuzzi. 2006. Insetos como indicadores de conservação da paisagem. Páginas 357-384, Capítulo 15 In: Biologia da Conservação: Essências. Rocha, C. F. D., H. G. Bergallo, M. Van Sluys &. M. A. S. Alves (editores). RiMa Editora, São Carlos., 587pp. (BOOK CHAPTER)

Andrade, R. B. & A. V. L. Freitas. 2005. Population biology of two species of Heliconius (Nymphalidae: Heliconiinae) in a semi-deciduous forest in Southeastern Brazil. Journal of the Lepidopterists’ Society 59(4): 223-228.

Francini, R. B., A. V. L. Freitas & K. S. Brown Jr. 2005. Rediscovery of Actinote zikani (D’Almeida) (Nymphalidae, Heliconiinae, Acraeini): Natural history, population biology and conservation of an endangered butterfly in SE Brazil. Journal of the Lepidopterists’ Society 59(3): 134-142.

Freitas, A. V. L. 2005. Borboletas. Borboletas da Mata Atlântica. Publicação privada (1000 cópias) – Tuim Parque. 12 pp.

Lewinsohn, T. M., A. V. L. Freitas & P. I. Prado. 2005. Conservation of Terrestrial Invertebrates and Their Habitats in Brazil. Conservation Biology, 19(3): 640-645.

Lewinsohn, T. M., A. V. L. Freitas & P. I. Prado. 2005. Conservação de Invertebrados Terrestres e Seus Hábitats no Brasil. Megadiversidade 1 (1): 62-69.

Uehara-Prado, M., A. V. L. Freitas & K. S. Brown Jr. 2005. Biological traits of frugivorous butterflies in a fragmented and a continuous landscape in the South Brazilian Atlantic Forest. Journal of the Lepidopterists’ Society 59(2): 96-106.

Oliveira, P. S. & A. V. L. Freitas. 2004. Ant-Plant-Herbivore Interactions in the Neotropical Cerrado Savanna. Review. Naturwissenschaften, 91(12): 557-570.

Schulz, S., G. Beccaloni, K. S. Brown Jr., M. Boppré, A. V. L. Freitas, P. Ockenfels & J. R. Trigo. 2004. Semiochemicals derived from pyrrolizidine alkaloids in male ithomiine butterflies (Lepidoptera: Nymphalidae: Ithomiinae). Biochemical Systematics and Ecology, 32 (8): 699-713.

Brown Jr., K. S. & A. V. L. Freitas. 2003. Butterfly Communities of Urban Forest Fragments in Campinas, São Paulo, Brazil: Structure, Instability, Environmental Correlates, and Conservation. Journal of Insect Conservation, 6(4): 217-231.

Freitas, A. V. L., R. B. Francini & K. S. Brown Jr. 2003. Insetos como indicadores ambientais. Páginas 125-151, Capítulo 5 In: Métodos de estudos em biologia da conservação e manejo da vida silvestre. Cullen Jr., L., R. Rudran & C. Valladares-Pádua (organizadores). Editora da UFPR; Fundação O Boticário de Proteção à Natureza, 667pp. (BOOK CHAPTER)

Raimundo, R. L. G., A. V. L. Freitas, R. N. S. Costa, J. B. F. Oliveira, A. B. Melo & K. S. Brown Jr. 2003. Manual de monitoramento ambiental usando borboletas e libélulas – Reserva Extrativista do Alto Juruá, 36 pp. (il). Série Pesquisa e Monitoramento Participativo em Áreas de Conservação Gerenciadas por Populações Tradicionais, Volume 1. Campinas, CERES/Laboratório de Antropologia e Ambiente. ISBN: 85-86572-13-6 (BOOK)

Brown Jr., K. S. & A. V. L. Freitas. 2002. Diversidade Biológica no Alto Juruá: Avaliação, Causas e Manutenção. Páginas 33-42; pranchas 13-16 In: Enciclopédia da floresta. O Alto Juruá: Práticas e conhecimentos das populações. Carneiro da Cunha, M. M. & M. B. Almeida (organizadores). São Paulo: Companhia das Letras, 735pp. (BOOK CHAPTER)

Oliveira, P. S., A. V. L. Freitas & K. Del-Claro. 2002. Ant foraging on plant foliage: contrasting effects on the behavioral ecology of insect herbivores. Páginas 287-305 In: The Cerrados of Brazil: Ecology and Natural History of a Neotropical Savanna. Oliveira, P. S. & R. J. Marquis (editores). Columbia University Press, NY, viii + 398pp. (BOOK CHAPTER)

Freitas, A. V. L., J. Vasconcellos-Neto, F. Vanini, J. R. Trigo & K. S. Brown Jr. 2001. Population studies of Aeria olena and Tithorea harmonia (Nymphalidae, Ithomiinae) in Southeastern Brazil. Journal of the Lepidopterists’ Society 55(4): 150-157.

Freitas, A. V. L. & R. R. Ramos. 2001. Population biology of Parides anchises nephalion (Papilionidae) in a coastal site in Southeast Brazil. Brazilian Journal of Biology, 61(4): 623-630.

Machado, G. & A. V. L. Freitas. 2001. Larval defence against ant predation in the butterfly Smyrna blomfildia. Ecological Entomology, 26: 436-439.

Brown Jr., K. S. & A. V. L. Freitas. 2000. Diversidade de Lepidoptera em Santa Teresa, Espírito Santo. Boletim do Museu de Biologia Mello Leitão, Nova Série, 11/12: 71-116.

Brown Jr., K. S. & A. V. L. Freitas. 2000. Atlantic Forest Butterflies: Indicators for Landscape Conservation. Biotropica, 32 (4b): 934-956.


Metodologia

Freitas, A. V. L., J. P. Santos, A. H. B. Rosa, C. A. Iserhard, A. Richter, R. R. Siewert, P. E. Gueratto, J. Y. O. Carreira & G. M. Lourenço. 2021. Sampling Methods for Butterflies (Lepidoptera). pp 101-123. In Santos, J. C. & G. W. Fernandes (eds). Measuring Arthropod Biodiversity. Springer, Cham. [PDF]

Rosa, A. H. B., L. V. Perillo, F. S. Neves, D. B. Ribeiro & A. V. L. Freitas. 2019. Butterflies collected using malaise traps as useful bycatches for ecology and conservation. Journal of Threatened Taxa 11(9): 14235-14237. [PDF]

Freitas, A. V. L., C. A. Iserhard, J. P. Santos, J. Y. O. Carreira, D. B. Ribeiro, D. H. A. melo, A. H. B. Rosa, O. J. Marini-Filho, G. M. Accacio & M. Uehara-Prado. 2014. Studies with butterfly bait traps: an overview. Revista Colombiana de Entomología 40 (2): 209-218. [PDF]

Silva-Brandão, K. L., M. L. Lyra & A. V. L. Freitas. 2009. Barcoding Lepidoptera: Current situation and perspectives on the usefulness of a contentious technique. Neotropical Entomology 38(4): 441-451.


Listas, Catálogos & Guias de identificação

Mota, L. L., S. J. Boddington, K. S. Brown Jr., C. J. Callaghan, G. Carter, W. Carter, S. M. Dantas, D. R. Dolibaina, K. Garwood, R. C. Hoyer, R. K. Robbins, A. Soh, K. R. Willmott & A. V. L. Freitas. 2022. The butterflies of Cristalino Lodge, in the Brazilian southern Amazonia: An updated species list with a significant contribution from citizen science. Biota Neotropica 22(3): e20221367. [PDF] [Vídeo]

Silva, L. D., A. H. B. Rosa, A. V. L. Freitas, C. A. Iserhard, L. A. Kaminski, O. J. Marini-Filho, B. M. Thompson, G. R. Soares & M. M. de Sousa. 2022. Papilionoidea inventory of the Sempre Vivas National Park, Minas Gerais State, southeastern Brazil (Insecta: Lepidoptera). SHILAP Revista de Lepidopterología 50: 367-380. [PDF]

Silva-Neto, A.M., Falaschi, R.L., Zacca, T., Hipólito, J., Pequeno, P.A.C.L., Alves-Oliveira, J.R., Santos, R.O. dos, Heleodoro, R.A., Jacobina, A.C. M., Somavilla, A., Camargo , A., Lira, A. de O., Sampaio , A.A., Ferreira, A. da S., Martins, A.L., Oliveira, A.F. de, Wengrat, A.P.G. da S., Rosa, A.H.B., Corrêa, C.C.D., De-Souza, C. C., Anjos-Santos, D., Cordeiro, D.P., Nogueira, D.S., Marques, D.W.A., Barbosa, D.N., Mendes, D.M. de M., Pádua, D.G. de, Vilela, D.S., Viegas, E.F.G., Santos , E.C. dos, Dias, F.M.S., Nascimento, F.E. de L., Duque, F.J.S., Mendes, G.C., Dantas, G.P. da S., Desidério, G.R., Marques, G.A., Menezes, I.S., Rodrigues, H.D.D., Fogaça, J.M., Schoeninger, K., Queiroz, L.L. de, Santana, L., Migliore, L.J., Fusari , L.M., Barros, L.M., Araújo, M.X., Cutrim, M., de Santis, M.D., Savaris, M., Gottschalk, M.S., Aragão, M., Bevilaqua, M., Bento, M., Soares, M.M.M., Reategui, N.S., Cipola, N.G., Saguiah, P.M., Pinto, P.J.C., Riccardi, P.R., Bartholomay, P.R., Boldrini, R., Sousa, R., Sobral , R., Mello , R.L., Carrenho, R., Olivier, R. da S., Siewert, R.R., Araujo, R. de O., Brito, R., Gadelha, S. de S., Moraes, S. de S., Pereira, T.P.L., Mahlmann, T. and Fernandes, D.R.R. 2022. Hexapoda Yearbook (Arthropoda: Mandibulata: Pancrustacea) Brazil 2020: the first annual production survey of new Brazilian species. EntomoBrasilis 15: e1000. [PDF]

L. T. Shirai & M. A. Stanton. 2021. Catálogo de borboletas do Parque Estadual Intervales, São Paulo, Brasil. Zenodo. Disponível em http://doi.org/10.5281/zenodo.5068939

Rodríguez-Ramírez, J. V., S. D. Ríos, S. S. Moraes, F. C. Penco & J. M. González. 2020. Catálogo de los Castniidae (Lepidoptera) del Museo Argentino de Ciencias Naturales “Bernardino Rivadavia”, Buenos Aires: descripción de tipos y notas generales e históricas. Revista del Museo Argentino de Ciencias Naturales 22(2): 173-190. [PDF]

Melo, D. H. A., M. Duarte, O. H. H. Mielke, R. K. Robbins & A. V. L. Freitas. 2019. Butterflies (Lepidoptera: Papilionoidea) of an urban park in northeastern Brazil. Biota Neotropica 19(1): e20180614.

Shirai, L. T., P. A. Machado, L. L. Mota, A. H. B. Rosa & A. V. L. Freitas. 2019. DnB, the Database of nymphalids in Brazil, with a checklist for standardized species lists. Journal of the Lepidopterists’ Society 73(2): 93-108.

Shirai, L. T., L. L. Mota, & A. V. L. Freitas. 2019. A treasure hunt for mistakes in DnB, the database of nymphalids of Brazil. News of the Lepidopterist’s Society 61: 136-138. [PDF]

Dos Santos, J. P., A. V. L. Freitas, K. S. Brown, J. Y. O. Carreira, P. E. Gueratto, A. H. B. Rosa, G. M. Lourenço, G. M. Accacio, M. Uehara-Prado, C. A. Iserhard, A. Richter, K. Gawlinski, H. P. Romanowski, N. O. Mega, M. O. Teixeira, A. Moser, D. B. Ribeiro, P. F. Araujo, B. K. C. Filgueiras, D. H. A. Melo, I. R. Leal, M. V. Beirão, S. P. Ribeiro, E. C. B. Cambuí, R. N. de Vasconcelos, Márcio Zikán Cardoso, Marlon Paluch, R. R. Greve, J. C. Voltolini, M. Galetti, A. L. Regolin, T. Sobral-Souza, M. C. Ribeiro. 2018. Atlantic butterflies: a data set of fruit-feeding butterfly communities from the Atlantic forests. Ecology 99: 2875-2875. [PDF]

Freitas, A. V. L., K. S. Brown Jr., O. H. H. Mielke, J. P. Santos & J. Vasconcellos-Neto. 2016. Borboletas da Reserva Natural Vale, Linhares/ES. Páginas 317-328. Capítulo 19. In: Floresta Atlântica de Tabuleiro: Diversidade e Endemismos na Reserva Natural Vale. Rolim, S. G., L. F. T. Menezes & A. C. Srbek-Araujo (Editores). Editora Rupestre. 496 pp.

Santos, J. P., A. V. L. Freitas, P. A. L. Constantino & M. Uehara-Prado. 2014. Guia de identificação de tribos de borboletas frugívoras. Amazônia. In. Pereira, A. B. & P. A. L. Constantino (Eds.). Monitoramento de Biodiversidade. MMA/ICMBio/GIZ. 12 pp.

Santos, J. P., A. V. L. Freitas, P. A. L. Constantino & M. Uehara-Prado. 2014. Guia de identificação de tribos de borboletas frugívoras. Cerrado. In. Pereira, A. B. & P. A. L. Constantino (Eds.). Monitoramento de Biodiversidade. MMA/ICMBio/GIZ. 12 pp.

Santos, J. P., A. V. L. Freitas, P. A. L. Constantino & M. Uehara-Prado. 2014. Guia de identificação de tribos de borboletas frugívoras. Mata Atlântica – Norte. In. Pereira, A. B. & P. A. L. Constantino (Eds.). Monitoramento de Biodiversidade. MMA/ICMBio/GIZ. 12 pp.

Santos, J. P., A. V. L. Freitas, P. A. L. Constantino & M. Uehara-Prado. 2014. Guia de identificação de tribos de borboletas frugívoras. Mata Atlântica – Sul. In. Pereira, A. B. & P. A. L. Constantino (Eds.). Monitoramento de Biodiversidade. MMA/ICMBio/GIZ. 12 pp.

Paluch, M., O. H. H. Mielke, C. E. B. Nobre, M. M. Casagrande, D. H. A. Melo & A. V. L. Freitas. 2012. Butterflies (Lepidoptera: Papilionoidea and Hesperioides) of the Parque Ecológico João Vasconcelos Sobrinho, Caruaru, Pernambuco, Brazil. Biota Neotropica, 11(4): 229-238.

Francini, R. B., M. Duarte, O. H. H. Mielke, A. Caldas & A. V. L. Freitas. 2011. Butterflies (Lepidoptera, Papilionoidea and Hesperioidea) of the “Baixada Santista” region, coastal São Paulo, Southeastern Brazil. Revista Brasileira de Entomologia, 55(1): 55-68.

Freitas, A. V. L. & O. J. Marini-Filho (Organizadores). 2011. Plano de ação nacional para a conservação dos lepidópteros ameaçados de extinção. Série Espécies Ameaçadas nº 13. ICMBio, Brasília, Brasil. 124p. (BOOK) [PDF]

Freitas, A. V. L., A. Camargo, C. A. Iserhard, G. M. Accacio, H. P. Romanowski, J. Galinkin, K. S. Brown Jr, M. Duarte, M. Uehara-Prado, M. Z. Cardoso, M. M. Casagrande, O. H. H. Mielke, O. J. Marini Filho, R. F. Monteiro, R. W. H. Honegger & W. L. Overal. 2011. Parte I. Informações Gerais. Páginas 15-74. In: Plano de ação nacional para a conservação dos lepidópteros ameaçados de extinção. Série Espécies Ameaçadas nº 13. Freitas, A. V. L. & O. J. Marini-Filho (Organizadores). ICMBio, Brasília, Brasil. 124p. (BOOK CHAPTER)

Duarte, M., R. K. Robbins, A. V. L. Freitas, K. S. Brown Jr., R. F. Monteiro, M. M. Casagrande, O. H. H. Mielke, M. S. Nascimento & T. G. Alves. 2010. Borboletas da Mata Atlântica do Estado do Rio de Janeiro: Lycaenidae (Lepidoptera). Arquivos do Museu Nacional do Rio de Janeiro, 67 (3-4): 291-302.

Monteiro, R. F., A. V. L. Freitas, M. A. F. Costa-Filho, M. S. Nascimento, T. G. Alves, K. S. Brown Jr., O. H. H. Mielke, M. M. Casagrande & M. Duarte. 2010. Borboletas da Mata Atlântica do Estado do Rio de Janeiro: Pieridae (Lepidoptera). Arquivos do Museu Nacional do Rio de Janeiro, 67 (3-4): 283-289.

Uehara-Prado, M., A. V. L. Freitas, R. B. Francini & K. S. Brown Jr. 2004. Guia das borboletas frugívoras da Reserva Estadual do Morro Grande e região de Caucaia do Alto, Cotia (SP). Biota Neotropica, 4(1): 1-25.


Borboletas frugívoras

Melo, D. H. A., A. V. L. Freitas, M. Tabarelli, B. K. C. Filgueiras & I. R. Leal. 2022. Aridity and chronic anthropogenic disturbance as organizing forces of fruit-feeding butterfly assemblages in a Caatinga dry forest. Biotropica, 00, 1–12. [PDF]

Álvares, C. F., A. Clavijo-Giraldo, S. I. Uribe, T. W. Pyrcz, C. A. Iserhard, A. V. L. Freitas & M. A. Marín. 2022. Sampling performance of bait traps in high Andean fruit-feeding butterflies. Neotropical Biodiversity 7(1): 507-513. [PDF]

Lourenço, G. M., W. Dáttilo, S. P. Ribeiro & A. V. L. Freitas. 2021. Biological Aspects and Movements of Neotropical Fruit-Feeding Butterflies. Neotropical Entomology 51: 43-53. [PDF]

Araujo, P. F., A. V. L. Freitas, G. A. S. Gonçalvez & D. B. Ribeiro. 2021. Vertical stratification on a small scale: the distribution of fruit-feeding butterflies in a semi-deciduous Atlantic forest in Brazil. Studies on Neotropical Fauna and Environment 56(1): 10-39. [PDF]

Freitas, A. V. L., D. G. Muniz, J. Y. O. Carreira & P. S. Oliveira. 2021. Fruit-Feeding Butterfly Assemblages in a Neotropical Savanna: Assessing Phenological Patterns Using Baited Traps. The Journal of the Lepidopterists’ Society 75(2): 88-103. [PDF]

Sobral-Souza, T., J. Stropp, J. P. Santos, V. M. Prasniewski, N. Szinwelski, B. Vilela, A. V. L. Freitas, M. C. Ribeiro & J. Hortal. 2021. Knowledge gaps hamper understanding the relationship between fragmentation and biodiversity loss: the case of Atlantic Forest fruit-feeding butterflies. PeerJ 9:e11673. [PDF]

Freire, G. B., T. Silva, H. Oliveira, C. Collier, H. P. Rodrigues, J. P. Dias, J. P. Santos, O. J. Marini-Filho, A. V. L. Freitas, A. M. Smilanich, L. A. Dyer & Ivone R. Diniz. 2021. Good Things Come in Larger Packages: Size Matters for Adult Fruit-Feeding Butterfly Dispersal and Larval Diet Breadth. Diversity 13(12): 664. [PDF]

Santos, J. P., T. Sobral-Souza, K. S. Brown Jr., M. H. Vancine, M. C. Ribeiro & A. V. L. Freitas. 2020. Effects of landscape modification on species richness patterns of fruit-feeding butterflies in Brazilian Atlantic Forest. Diversity and Distributions 26(2): 196-208.

Gueratto, P. E., J. Y. O. Carreira, J. P. Santos, A. Tacioli & A. V. L. Freitas. 2020. Effects of forest trails on the community structure of tropical butterflies. Journal of Insect Conservation 24(2): 309-319. [PDF]

Lourenço, G. M., P. Luna, R. Guevara, W. Dattilo, A. V. L. Freitas & S. P. Ribeiro. 2020. Temporal shifts in butterfly diversity: responses to natural and anthropic forest transitions. Journal of Insect Conservation 24(2): 353-363. [PDF]

Dos Santos, J. P., A. V. L. Freitas, K. S. Brown, J. Y. O. Carreira, P. E. Gueratto, A. H. B. Rosa, G. M. Lourenço, G. M. Accacio, M. Uehara-Prado, C. A. Iserhard, A. Richter, K. Gawlinski, H. P. Romanowski, N. O. Mega, M. O. Teixeira, A. Moser, D. B. Ribeiro, P. F. Araujo, B. K. C. Filgueiras, D. H. A. Melo, I. R. Leal, M. V. Beirão, S. P. Ribeiro, E. C. B. Cambuí, R. N. de Vasconcelos, Márcio Zikán Cardoso, Marlon Paluch, R. R. Greve, J. C. Voltolini, M. Galetti, A. L. Regolin, T. Sobral-Souza, M. C. Ribeiro. 2019. Fruit-Feeding Butterflies from the Atlantic Forests. The Bulletin of the Ecological Society of America (Online) 100: p. e01484. [PDF]

Filgueiras, B. K. C., D. H. A. Melo, M. Uehara-Prado, A. V. L. Freitas, I. R. Leal & M. Tabarelli. 2019. Compensatory dynamics on the community structure of fruit-feeding butterflies across hyper-fragmented Atlantic forest habitats. Ecological Indicators 98: 276-284.

Lourenço, G. M., G. R. Soares, T. P. Santos, W. Dattilo, A. V. L. Freitas & S. P. Ribeiro. 2019. Equal but different: Natural ecotones are dissimilar to anthropic edges PLoS ONE 14(3): e0213008. 18pp. [PDF]

Melo, D. H. A., B. K. C. Filgueiras, C. A. Iserhard, L. Iannuzzi, A. V. L. Freitas & I. R. Leal. 2019. Effect of habitat loss and fragmentation on fruit-feeding butterflies in the Brazilian Atlantic Forest. Canadian Journal of Zoology, 97(7): 588-596. [PDF]

Dos Santos, J. P., A. V. L. Freitas, K. S. Brown, J. Y. O. Carreira, P. E. Gueratto, A. H. B. Rosa, G. M. Lourenço, G. M. Accacio, M. Uehara-Prado, C. A. Iserhard, A. Richter, K. Gawlinski, H. P. Romanowski, N. O. Mega, M. O. Teixeira, A. Moser, D. B. Ribeiro, P. F. Araujo, B. K. C. Filgueiras, D. H. A. Melo, I. R. Leal, M. V. Beirão, S. P. Ribeiro, E. C. B. Cambuí, R. N. de Vasconcelos, Márcio Zikán Cardoso, Marlon Paluch, R. R. Greve, J. C. Voltolini, M. Galetti, A. L. Regolin, T. Sobral-Souza, M. C. Ribeiro. 2019. Fruit-Feeding Butterflies from the Atlantic Forests. The Bulletin of the Ecological Society of America (Online) 100: p. e01484. [PDF]

Dos Santos, J. P., A. V. L. Freitas, K. S. Brown, J. Y. O. Carreira, P. E. Gueratto, A. H. B. Rosa, G. M. Lourenço, G. M. Accacio, M. Uehara-Prado, C. A. Iserhard, A. Richter, K. Gawlinski, H. P. Romanowski, N. O. Mega, M. O. Teixeira, A. Moser, D. B. Ribeiro, P. F. Araujo, B. K. C. Filgueiras, D. H. A. Melo, I. R. Leal, M. V. Beirão, S. P. Ribeiro, E. C. B. Cambuí, R. N. de Vasconcelos, Márcio Zikán Cardoso, Marlon Paluch, R. R. Greve, J. C. Voltolini, M. Galetti, A. L. Regolin, T. Sobral-Souza, M. C. Ribeiro. 2018. Atlantic butterflies: a data set of fruit-feeding butterfly communities from the Atlantic forests. Ecology 99: 2875-2875. [PDF]

de Andrade, R. B., J. K. Balch, J. Y. O. Carreira, P. M. Brando & A. V. L. Freitas. 2017. The impacts of recurrent fires on diversity of fruit-feeding butterflies in a south-eastern Amazon forest. Journal of Tropical Ecology 33 (1): 22-32. [PDF]

Santos, J. P., C. A. Iserhard, J. Y. O. Carreira & A. V. L. Freitas. 2017. Monitoring fruit-feeding butterfly assemblages in two vertical strata in seasonal Atlantic Forest: temporal species turnover is lower in the canopy. Journal of Tropical Ecology 33(5): 345-355. [PDF]

Filgueiras, B. K. C., D. H. A. Melo, I. R. Leal, M. Tabarelli, A. V. L. Freitas & Luciana Iannuzzi. 2016. Fruit-feeding butterflies in edge-dominated habitats: community structure, species persistence and cascade effect. Journal of Insect Conservation 20(3): 539-548.

Freitas, A. V. L., C. A. Iserhard, J. P. Santos, J. Y. O. Carreira, D. B. Ribeiro, D. H. A. melo, A. H. B. Rosa, O. J. Marini-Filho, G. M. Accacio & M. Uehara-Prado. 2014. Studies with butterfly bait traps: an overview. Revista Colombiana de Entomología 40 (2): 209-218. [PDF]

Sant’Anna, C. L. B., D. B. Ribeiro, L. C. Garcia & A. V. L. Freitas. 2014. Fruit-feeding butterfly communities are influenced by restoration age in tropical forests. Restoration Ecology 22(4): 480-485.

Santos, J. P., A. V. L. Freitas, P. A. L. Constantino & M. Uehara-Prado. 2014. Guia de identificação de tribos de borboletas frugívoras. Amazônia. In. Pereira, A. B. & P. A. L. Constantino (Eds.). Monitoramento de Biodiversidade. MMA/ICMBio/GIZ. 12 pp.

Santos, J. P., A. V. L. Freitas, P. A. L. Constantino & M. Uehara-Prado. 2014. Guia de identificação de tribos de borboletas frugívoras. Cerrado. In. Pereira, A. B. & P. A. L. Constantino (Eds.). Monitoramento de Biodiversidade. MMA/ICMBio/GIZ. 12 pp.

Santos, J. P., A. V. L. Freitas, P. A. L. Constantino & M. Uehara-Prado. 2014. Guia de identificação de tribos de borboletas frugívoras. Mata Atlântica – Norte. In. Pereira, A. B. & P. A. L. Constantino (Eds.). Monitoramento de Biodiversidade. MMA/ICMBio/GIZ. 12 pp.

Santos, J. P., A. V. L. Freitas, P. A. L. Constantino & M. Uehara-Prado. 2014. Guia de identificação de tribos de borboletas frugívoras. Mata Atlântica – Sul. In. Pereira, A. B. & P. A. L. Constantino (Eds.). Monitoramento de Biodiversidade. MMA/ICMBio/GIZ. 12 pp.

Ribeiro, D. B. & A. V. L. Freitas. 2012. The effect of reduced-impact logging on fruit-feeding butterflies in Central Amazon, Brazil. Journal of Insect Conservation, 16: 733-744.

Ribeiro, D. B., R. Batista, P. I. Prado, K. S. Brown Jr. & A. V. L. Freitas. 2012. The importance of small scales to the fruit-feeding butterfly assemblages in a fragmented landscape. Biodiversity and Conservation, 21: 811-827.

Ribeiro, D. B. & A. V. L. Freitas. 2011. Large-sized insects show stronger seasonality than small-sized ones: a case study of fruit-feeding butterflies. Biological Journal of the Linnean Society, (104): 820–827.

Ribeiro, D. B. & A. V. L. Freitas. 2010. Differences in thermal responses in a fragmented landscape: Temperature affects the sampling of diurnal, but not nocturnal fruit-feeding Lepidoptera. The Journal of Research on the Lepidoptera 42: 1-4.

Ribeiro, D. B., P. I. Prado, K. S. Brown Jr. & A. V. L. Freitas. 2010. Temporal diversity patterns and phenology in fruit-feeding butterflies in the Atlantic Forest. Biotropica, 42(6): 710-716.

Uehara-Prado, M., J. O. Fernandes, A. M. Bello, G. Machado, A. J. Santos, F. Z. Vaz-de-Mello & A. V. L. Freitas. 2009. Selecting terrestrial arthropods as indicators of small-scale disturbance: a first approach in the Brazilian Atlantic Forest. Biological Conservation 142: 1220-1228.

Ribeiro, D. B., P. I. Prado, K. S. Brown Jr. & A. V. L. Freitas. 2008. Additive partitioning of butterfly diversity in a fragmented landscape: Importance of scale and implications for conservation. Diversity and Distributions, 14: 961-968.

Brown Jr K. S., A. V. L. Freitas, B. Von Schoultz, A. O. Saura & A. Saura. 2007. Chromosomal evolution of South American frugivorous butterflies in the Satyroid clade (Nymphalidae: Charaxinae, Morphinae and Satyrinae). Biological Journal of the Linnean Society, 92: 467-481.

Uehara-Prado, M. K. S. Brown Jr. & A. V. L. Freitas. 2007. Species richness, composition and abundance of fruit-feeding butterflies in the Brazilian Atlantic Forest: comparison between a fragmented and a continuous landscape. Global Ecology and Biogeography, 16: 43-54.

Uehara-Prado, M., A. V. L. Freitas & K. S. Brown Jr. 2005. Biological traits of frugivorous butterflies in a fragmented and a continuous landscape in the South Brazilian Atlantic Forest. Journal of the Lepidopterists’ Society 59(2): 96-106.

Uehara-Prado, M., A. V. L. Freitas, R. B. Francini & K. S. Brown Jr. 2004. Guia das borboletas frugívoras da Reserva Estadual do Morro Grande e região de Caucaia do Alto, Cotia (SP). Biota Neotropica, 4(1): 1-25.


Mariposas

Machado, P. A., P. E. Gueratto, S. S. Moraes, A. R. Nascimento, M. J. Kato & A. V. L. Freitas. 2022. Temporal Distribution of Eois Immatures (Lepidoptera, Geometridae) in a Tropical Forest. The Journal of the Lepidopterists’ Society 76(4): 261-267. [PDF]

Moraes, S. S., P. E. Gueratto, J. P. dos Santos, M. H. Santos, A. V. L. Freitas & M. Duarte. 2022. Niche modelling and comparative morphology untangle taxonomy of the Dysschema eurocilia clade (Lepidoptera: Erebidae) and reveal a relictual Pleistocene Arc distribution. Systematics and Biodiversity 20:1. [PDF]

Moraes, S. S., L. Murillo-Ramos, P. A. Machado, H. R. Ghanavi, L. Magaldi, K. L. Silva-Brandão, M. J. Kato, A. V. L. Freitas & N. Wahlberg. 2021. A Double-Edged Sword: Unrecognized cryptic diversity and taxonomic impediment in Eois (Lepidoptera: Geometridae). Zoologica Scripta 2021: 1-14. [PDF]

Moraes, S. S., Y. Montebello, M. A. Stanton, L. F. Yamaguchi, M. J. Kato & A. V. L. Freitas. 2021. Description of three new species of Geometridae (Lepidoptera) using species delimitation in an integrative taxonomy approach for a cryptic species complex. PeerJ 9: e11304. [PDF]

Rodríguez-Ramírez, J. V., S. D. Ríos, S. S. Moraes, F. C. Penco & J. M. González. 2020. Catálogo de los Castniidae (Lepidoptera) del Museo Argentino de Ciencias Naturales “Bernardino Rivadavia”, Buenos Aires: descripción de tipos y notas generales e históricas. Revista del Museo Argentino de Ciencias Naturales 22(2): 173-190. [PDF]

Cosmo, L. G.A. R. Nascimento, R. Cogni & A. V. L. Freitas. 2019. Temporal distribution in a tri-trophic system associated with Piper amalago L. in a tropical seasonal forest. Arthropod-Plant Interactions 13(4): 647-652. [PDF]

Freitas, A. V. L. 2017. Biological notes on Utetheisa pulchelloides (Erebidae) in Tuamotu, French Polynesia. Tropical Lepidoptera Research 27(2): 115-116.

Zenker, M. M., N. Wahlberg, G. Brehm, J. A. Teston, L. Przybylowicz, M. R. Pie & A. V. L. Freitas. 2017. Systematics and origin of moths in the subfamily Arctiinae (Lepidoptera, Erebidae) in the Neotropical region. Zoologica Scripta 46(3): 348-362.

Zenker, M. M., A. V. L. Freitas, R. Rougerie, M. Laguerre, J. A. Teston & M. R. Pie. 2016. Uncovering tiger moth diversity in Brazil’s most endangered biome. Barcode Bulletin 7(2): 22-23.

Zenker, M. M., R. Rougerie, J. A. Teston, M. Laguerre, M. R. Pie & A. V. L. Freitas. 2016. Fast census of moth diversity in the Neotropics: A comparison of field-assigned morphospecies and DNA barcoding in tiger moths. PLoS ONE 11(2): e0148423.

Zenker, M. M., P. J. DeVries, C. M. Penz, J. A. Teston, A. V. L. Freitas & M. R. Pie. 2015. Diversity and composition of Arctiinae moth assemblages along elevational and spatial dimensions in Brazilian Atlantic Forest. Journal of Insect Conservation 19(1): 129–140.

Ribeiro, D. B. & A. V. L. Freitas. 2010. Differences in thermal responses in a fragmented landscape: Temperature affects the sampling of diurnal, but not nocturnal fruit-feeding Lepidoptera. The Journal of Research on the Lepidoptera 42: 1-4.

Algarin, C. M., M. A. Marin, S. Uribe & A. V. L. Freitas. 2008. Tabla de vida de Pseudodirphia pallida (Lepidoptera: Saturniidae) en condiciones de laboratório. Revista Colombiana de Entomologia, 34(1): 116-120.


Mata Atlântica

Machado, P. A., P. E. Gueratto, S. S. Moraes, A. R. Nascimento, M. J. Kato & A. V. L. Freitas. 2022. Temporal Distribution of Eois Immatures (Lepidoptera, Geometridae) in a Tropical Forest. The Journal of the Lepidopterists’ Society 76(4): 261-267. [PDF]

Rosa, A. H. B., N. Seraphim, P. A. Machado, P. E. Gueratto, T. Sobral-Souza & A. V. L. Freitas. 2022. Eucorna sanarita (Schaus) (Riodinidae: Riodininae): Distribution, systematic position, and conservation of a threatened Brazilian butterfly in the Atlantic Forest biodiversity hotspot. Journal of Insect Conservation (online). [PDF]

Shirai, L. T., M. A. Stanton, G. B. D’Angelo, T. B. Conforti, A. V. L. Freitas, M. J. Kato & L. F. Yamaguchi. 2022. Interaction Gardens and Butterfly Catalogues: a Joint Strategy to Promote Capacity Development in Protected Areas and Reduce the Extinction of Experience in Cities. Cities and the Environment (CATE) 15(1): article 3. [PDF]

Freitas, A. V. L. 2022. Immature stages of Splendeuptychia ambra (Nymphalidae: Euptychiina) and the diversity of immature morphology within Splendeuptychia. Tropical Lepidoptera Research 32(1): 52-58. [PDF]

Lourenço, G. M., W. Dáttilo, S. P. Ribeiro & A. V. L. Freitas. 2021. Biological Aspects and Movements of Neotropical Fruit-Feeding Butterflies. Neotropical Entomology 51: 43-53. [PDF]

Ramos, R.R., R. B. Francini, M. E. M. Habib & A. V. L. Freitas. 2021. Seasonal Patterns of Host Plant Use in an Assemblage of Heliconiini Butterflies (Lepidoptera: Nymphalidae) in a Neotropical forest. Neotropical Entomology. [PDF]

Pablos, J. L., A. K. Silva, N. Seraphim, L. M. Magaldi, A. P. Souza, A. V. L. Freitas & K. L. Silva-Brandão. 2021. North-south and climate-landscape-associated pattern of population structure for the Atlantic Forest White Morpho butterflies. Molecular Phylogenetics and Evolution 161: 107157. [PDF]

Freitas, A. V. L., A. H. B. Rosa, K. S. Brown Jr., C. E. B. Nobre, D. H. A. Melo & S. M. Kerpel. 2021. A new and rare Ithomiini from Northeastern Brazil (Lepidoptera: Nymphalidae: Danainae). Journal of the Lepidopterists’ Society 75(1): 44–48. [PDF]

A. V. L. Freitas, E. P. Barbosa & J. Y. O. Carreira. 2021. Immature stages and natural history of Yphthimoides borasta (Nymphalidae: Euptychiina). Tropical Lepidoptera Research 31(1): 42-47. [PDF]

A. V. L. Freitas, E. P. Barbosa & J. Y. O. Carreira. 2021. Immature stages and natural history of Yphthimoides borasta (Nymphalidae: Euptychiina). Tropical Lepidoptera Research 31(1): 42-47. [PDF]

Moraes, S. S., Y. Montebello, M. A. Stanton, L. F. Yamaguchi, M. J. Kato & A. V. L. Freitas. 2021. Description of three new species of Geometridae (Lepidoptera) using species delimitation in an integrative taxonomy approach for a cryptic species complex. PeerJ 9: e11304. [PDF]

Sobral-Souza, T., J. Stropp, J. P. Santos, V. M. Prasniewski, N. Szinwelski, B. Vilela, A. V. L. Freitas, M. C. Ribeiro & J. Hortal. 2021. Knowledge gaps hamper understanding the relationship between fragmentation and biodiversity loss: the case of Atlantic Forest fruit-feeding butterflies. PeerJ 9:e11673. [PDF]

L. T. Shirai & M. A. Stanton. 2021. Catálogo de borboletas do Parque Estadual Intervales, São Paulo, Brasil. Zenodo. Disponível em http://doi.org/10.5281/zenodo.5068939

Rosa, A. H. B., L. A. R. Leite, R. P. R. Costa & A. V. L. Freitas. 2021. A New Species of Dynamine Hübner, [1919] (Lepidoptera: Nymphalidae: Biblidinae) from the Atlantic Forest in Southeastern Brazil. Neotropical Entomology 50: 948–957. [PDF]

Santos, J. P., T. Sobral-Souza, K. S. Brown Jr., M. H. Vancine, M. C. Ribeiro & A. V. L. Freitas. 2020. Effects of landscape modification on species richness patterns of fruit-feeding butterflies in Brazilian Atlantic Forest. Diversity and Distributions 26(2): 196-208.

Gueratto, P. E., P. A. Machado, T. M. C. Aguiar, E. P. Barbosa, F. M. S. Dias, J. F. Oliveira-Neto, M. M. Casagrande & A. V. L. Freitas. 2020. Identifying Memphis: A comprehensive and comparative description of the immature stages and natural history of Memphis acidalia victoria (H. Druce, 1877) (Lepidoptera: Nymphalidae). Austral Entomology 59(1): 127-141. [PDF]

Gueratto, P. E., J. Y. O. Carreira, J. P. Santos, A. Tacioli & A. V. L. Freitas. 2020. Effects of forest trails on the community structure of tropical butterflies. Journal of Insect Conservation 24(2): 309-319. [PDF]

Lourenço, G. M., P. Luna, R. Guevara, W. Dattilo, A. V. L. Freitas & S. P. Ribeiro. 2020. Temporal shifts in butterfly diversity: responses to natural and anthropic forest transitions. Journal of Insect Conservation 24(2): 353-363. [PDF]

Freitas, A. V. L. 2020. A new clearwing butterfly from northeastern Brazil (Nymphalidae: Danainae: Ithomiini). Tropical Lepidoptera Research 30(1): 28-32. [PDF]

Barbosa, E. P., T. M. C. Aguiar, A. H. B. Rosa, T. Zacca & A. V. L. Freitas. 2020. A new species of butterfly (Lepidoptera: Nymphalidae) from the highlands of Southeastern Brazil. Zootaxa 4819(1): zootaxa.4819.1.6. [PDF]

Rosa, A. H. B., D. B. Ribeiro & A. V. L. Freitas. 2020. Population biology, natural history and conservation of two endangered high elevation Neotropical butterflies. Journal of Insect Conservation 24(4): 681-694. [PDF]

Brown, J. L., A. Paz, M. Reginato, C. A. Renata, C. Assis, M. Lyra, M. K. Caddah, J. Aguirre-Santoro, F. d’Horta, F. R. Amaral, R. Goldenberg, K. L. Silva-Brandão, A. V. L. Freitas, M. T. Rodrigues, F. A. Michelangeli, C. Y. Miyaki & A. C. Carnaval. 2020. Seeing the forest through many trees: Multi‐taxon patterns of phylogenetic diversity in the Atlantic Forest hotspot. Diversity and Distributions 26(9): 1160-1176. [PDF]

Freitas, A. V. L., R. B. Francini, O. H. H. Mielke, A. H. B. Rosa, L. M. Magaldi & K. L. Silva-Brandão. 2020. A New and Rare Actinote Hübner (Lepidoptera: Nymphalidae: Heliconiinae: Acraeini) from Southeastern Brazil. Neotropical Entomology 49: 696-703. [PDF]

Freitas, A. V. L., A. H. B. Rosa, C. E. B. Nobre, D. H. A. Melo, L. L. Mota, K. L. Silva-Brandão, P. A. Machado & J. Y. O. Carreira. 2020. Immature Stages, Natural History, Systematics and Conservation of an Endangered Neotropical Butterfly: the Case of Scada karschina delicata (Nymphalidae: Ithomiini). Neotropical Entomology 49(5): 685-695. [PDF]

Massardo, D., N. W. VanKuren, S. Nallu, R. R. Ramos, P. G. Ribeiro, K. L. Silva-Brandão, M. M. Brandão, M. B. Lion, A. V. L. Freitas, M. Z. Cardoso & M. R. Kronforst. The roles of hybridization and habitat fragmentation in the evolution of Brazil’s enigmatic longwing butterflies, Heliconius nattereri and H. hermathena. BMC Biology 18(1): 1-21. [PDF]

Filgueiras, B. K. C., D. H. A. Melo, M. Uehara-Prado, A. V. L. Freitas, I. R. Leal & M. Tabarelli. 2019. Compensatory dynamics on the community structure of fruit-feeding butterflies across hyper-fragmented Atlantic forest habitats. Ecological Indicators 98: 276-284.

Melo, D. H. A., M. Duarte, O. H. H. Mielke, R. K. Robbins & A. V. L. Freitas. 2019. Butterflies (Lepidoptera: Papilionoidea) of an urban park in northeastern Brazil. Biota Neotropica 19(1): e20180614.

Iserhard, C. A., L. Duarte, N. Seraphim & A. V. L. Freitas. 2019. How urbanization affects multiple dimensions of biodiversity in tropical butterfly assemblages. Biodiversity and Conservation 28(3): 621-638. [PDF]

Lourenço, G. M., G. R. Soares, T. P. Santos, W. Dattilo, A. V. L. Freitas & S. P. Ribeiro. 2019. Equal but different: Natural ecotones are dissimilar to anthropic edges PLoS ONE 14(3): e0213008. 18pp. [PDF]

Uehara-Prado, M & A. V. L. Freitas. 2019. Population structure of Taygetis ypthima (Nymphalidae, Euptychiina) in Southeast Brazil. Journal of the Lepidopterists’ Society 73(1): 63-64. [PDF]

Carvalho, M. R. M., E. P. Barbosa & A. V. L. Freitas. 2019. Immature stages of Mechanitis polymnia casabranca (Nymphalidae, Danainae). Tropical Lepidoptera Research 29(1): 12-18.

Freitas, A. V. L., M. S. Woodbury, E. P. Barbosa & K. R. Willmott. 2019. A new subspecies of Adelpha messana (C. Felder & R. Felder) from the Brazilian Atlantic forest (Nymphalidae: Limenitidinae). Tropical Lepidoptera Research 29(2): 62-66.

Melo, D. H. A., B. K. C. Filgueiras, C. A. Iserhard, L. Iannuzzi, A. V. L. Freitas & I. R. Leal. 2019. Effect of habitat loss and fragmentation on fruit-feeding butterflies in the Brazilian Atlantic Forest. Canadian Journal of Zoology, 97(7): 588-596. [PDF]

Cosmo, L. G.A. R. Nascimento, R. Cogni & A. V. L. Freitas. 2019. Temporal distribution in a tri-trophic system associated with Piper amalago L. in a tropical seasonal forest. Arthropod-Plant Interactions 13(4): 647-652. [PDF]

Dos Santos, J. P., A. V. L. Freitas, K. S. Brown, J. Y. O. Carreira, P. E. Gueratto, A. H. B. Rosa, G. M. Lourenço, G. M. Accacio, M. Uehara-Prado, C. A. Iserhard, A. Richter, K. Gawlinski, H. P. Romanowski, N. O. Mega, M. O. Teixeira, A. Moser, D. B. Ribeiro, P. F. Araujo, B. K. C. Filgueiras, D. H. A. Melo, I. R. Leal, M. V. Beirão, S. P. Ribeiro, E. C. B. Cambuí, R. N. de Vasconcelos, Márcio Zikán Cardoso, Marlon Paluch, R. R. Greve, J. C. Voltolini, M. Galetti, A. L. Regolin, T. Sobral-Souza, M. C. Ribeiro. 2019. Fruit-Feeding Butterflies from the Atlantic Forests. The Bulletin of the Ecological Society of America (Online) 100: p. e01484. [PDF]

Freitas A.V.L., Francini R.B., Paluch M., Barbosa E.P. 2018. A new species of Actinote Hübner (Nymphalidae: Heliconiinae: Acraeini) from southeast Brazil. Revista Brasileira de Entomologia 62: 135-147.

Pyrcz T. W., Freitas A. V. L., Boyer P., Dias F. M. S., Dolibaina D. R., Barbosa E. P., Magaldi L. M., Mielke O. H. H., Casagrande M. M. & Lorenc-Brudecka J. 2018. Uncovered Diversity of a Predominantly Andean Butterfly Clade in the Brazilian Atlantic Forest: a Revision of the Genus Praepedaliodes Forster (Lepidoptera: Nymphalidae, Satyrinae, Satyrini). Neotropical Entomology 47: 211–255. [PDF]

Freitas, A. V. L., A. H. B. Rosa, J. Y. O. Carreira, P. E. Gueratto, J. P. Santos, A. Tacioli. 2018. Immature stages of two Moneuptychia from southeastern Brazil (Nymphalidae: Euptychiina). Tropical Lepidoptera Research 28(2): 100-105. [PDF]

Dos Santos, J. P., A. V. L. Freitas, K. S. Brown, J. Y. O. Carreira, P. E. Gueratto, A. H. B. Rosa, G. M. Lourenço, G. M. Accacio, M. Uehara-Prado, C. A. Iserhard, A. Richter, K. Gawlinski, H. P. Romanowski, N. O. Mega, M. O. Teixeira, A. Moser, D. B. Ribeiro, P. F. Araujo, B. K. C. Filgueiras, D. H. A. Melo, I. R. Leal, M. V. Beirão, S. P. Ribeiro, E. C. B. Cambuí, R. N. de Vasconcelos, Márcio Zikán Cardoso, Marlon Paluch, R. R. Greve, J. C. Voltolini, M. Galetti, A. L. Regolin, T. Sobral-Souza, M. C. Ribeiro. 2018. Atlantic butterflies: a data set of fruit-feeding butterfly communities from the Atlantic forests. Ecology 99: 2875-2875. [PDF]

Nakahara, S., E. P. Barbosa & A. V. L. Freitas. 2017. A potentially endangered new species of Euptychia Hübner, 1818 (Lepidoptera: Nymphalidae: Satyrinae) from the Atlantic Coastal Forest of Brazil. Neotropical Entomology 46(3): 302-309.

Santos, J. P., C. A. Iserhard, J. Y. O. Carreira & A. V. L. Freitas. 2017. Monitoring fruit-feeding butterfly assemblages in two vertical strata in seasonal Atlantic Forest: temporal species turnover is lower in the canopy. Journal of Tropical Ecology 33(5): 345-355. [PDF]

Rosa, A. H. B., M. C. Campis, D. B. Ribeiro & A. V. L. Freitas. 2017. New record of the threatened butterfly Drephalys mourei (Hesperiidae) in a heavily disturbed area in Southeastern Brazil. Journal of the Lepidopterists’ Society 71(4): 278-280.

Shirai, L. T., L. L. Mota & A. V. L. Freitas. 2017. Aggregation of Epityches eupompe (Nymphalidae: Ithomiini) in southern Brazil. Tropical Lepidoptera Research 27(2): 111-114.

Freitas, A. V. L., J. Y. O. Carreira, J. P. Santos & E. P. Barbosa. 2016. Immature stages and natural history of two species of Forsterinaria from southeastern Brazil (Lepidoptera: Nymphalidae). Tropical Lepidoptera Research 26(1): 13-18. [PDF]

Freitas, A. V. L., K. S. Brown Jr., O. H. H. Mielke, J. P. Santos & J. Vasconcellos-Neto. 2016. Borboletas da Reserva Natural Vale, Linhares/ES. Páginas 317-328. Capítulo 19. In: Floresta Atlântica de Tabuleiro: Diversidade e Endemismos na Reserva Natural Vale. Rolim, S. G., L. F. T. Menezes & A. C. Srbek-Araujo (Editores). Editora Rupestre. 496 pp.

Freitas, A. V.L., T. M. Lewinsohn, L. M. Magaldi & N. Wahlberg. 2015. A new subspecies of Anthanassa (Nymphalidae: Nymphalinae: Melitaeini) from southeastern Brazil. Journal of the Lepidopterists’ Society 69(2): 83-90.

Zenker, M. M., P. J. DeVries, C. M. Penz, J. A. Teston, A. V. L. Freitas & M. R. Pie. 2015. Diversity and composition of Arctiinae moth assemblages along elevational and spatial dimensions in Brazilian Atlantic Forest. Journal of Insect Conservation 19(1): 129–140.

Melo, D. H. A., B. K. C. Filgueiras, I. R. Leal & A. V. L. Freitas. 2014. Recent records of Morpho menelaus eberti Fischer (Lepidoptera: Nymphalidae), an endangered butterfly in Northeast Brazil. Tropical Lepidoptera Research 24(2): 102-104.

Sant’Anna, C. L. B., D. B. Ribeiro, L. C. Garcia & A. V. L. Freitas. 2014. Fruit-feeding butterfly communities are influenced by restoration age in tropical forests. Restoration Ecology 22(4): 480-485.

Santos, J. P., A. V. L. Freitas, P. A. L. Constantino & M. Uehara-Prado. 2014. Guia de identificação de tribos de borboletas frugívoras. Mata Atlântica – Norte. In. Pereira, A. B. & P. A. L. Constantino (Eds.). Monitoramento de Biodiversidade. MMA/ICMBio/GIZ. 12 pp.

Santos, J. P., A. V. L. Freitas, P. A. L. Constantino & M. Uehara-Prado. 2014. Guia de identificação de tribos de borboletas frugívoras. Mata Atlântica – Sul. In. Pereira, A. B. & P. A. L. Constantino (Eds.). Monitoramento de Biodiversidade. MMA/ICMBio/GIZ. 12 pp.

Freitas, A. V. L., E. P. Barbosa, J. P. Santos & O. H. H. Mielke. 2013. A new genus, Atlanteuptychia gen. nov., for Euptychia ernestina (Lepidoptera: Nymphalidae: Satyrinae. Zoologia 30(6): 661-668.

Greve, R. R., C. Callaghan, L. A. Kaminski & A. V. L. Freitas. 2013. The rediscovery of Joiceya praeclarus Talbot 1928 (Lepidoptera: Riodinidae), more than 80 years after its description. Journal of the Lepidopterists’ Society, 67(1): 56-57.

Herkenhoff, E. V., R. F. Monteiro, A. P. Esperanço & A. V. L. Freitas. 2013. Population biology of the endangered fluminense swallowtail butterfly Parides ascanius (Papilionidae: Papilioninae: Troidini). Journal of the Lepidopterists’ Society, 67(1): 29-34.

Iserhard, C. A., K. S. Brown Jr. & A. V. L. Freitas. 2013. Maximized sampling of butterflies to detect temporal changes in tropical communities. Journal of Insect Conservation, 17: 615-622.

Freitas, A. V. L., L. A. Kaminski, O. H. H. Mielke, E. P. Barbosa & K. L. Silva-Brandão. 2012. A new species of Yphthimoides (Lepidoptera: Nymphalidae: Satyrinae) from the southern Atlantic forest region. Zootaxa, 3526: 31-44.

Ramos, R. R., D. Rodrigues & A. V. L. Freitas. 2012. Oviposition preference and larval performance in a Heliconius erato phyllis (Lepidoptera: Nymphalidae) population from southeastern Brazil: is there a positive relationship? Journal of Natural History, 46(11-12): 669-681.

Ribeiro, D. B., R. Batista, P. I. Prado, K. S. Brown Jr. & A. V. L. Freitas. 2012. The importance of small scales to the fruit-feeding butterfly assemblages in a fragmented landscape. Biodiversity and Conservation, 21: 811-827.

Francini, R. B., M. Duarte, O. H. H. Mielke, A. Caldas & A. V. L. Freitas. 2011. Butterflies (Lepidoptera, Papilionoidea and Hesperioidea) of the “Baixada Santista” region, coastal São Paulo, Southeastern Brazil. Revista Brasileira de Entomologia, 55(1): 55-68.

Francini, R. B., E. P. Barbosa & A. V. L. Freitas. 2011. Immature stages of Actinote zikani (Nymphalidae: Heliconiinae), a critically endangered butterfly from southeastern Brazil. Tropical Lepidoptera Research 21(1): 20-26.

Freitas, A. V. L., L. A. Kaminski, C. A. Iserhard, E. P. Barbosa & O. J. Marini-Filho. 2011. The endangered butterfly Charonias theano (Boisduval) (Lepidoptera: Pieridae): current status, threats and its rediscovery in São Paulo state, southeastern Brazil. Neotropical Entomology, 40(6): 669-676.

Ribeiro, D. B. & A. V. L. Freitas. 2011. Large-sized insects show stronger seasonality than small-sized ones: a case study of fruit-feeding butterflies. Biological Journal of the Linnean Society, (104): 820–827.

Monteiro, R. F., A. V. L. Freitas, M. A. F. Costa-Filho, M. S. Nascimento, T. G. Alves, K. S. Brown Jr., O. H. H. Mielke, M. M. Casagrande & M. Duarte. 2010. Borboletas da Mata Atlântica do Estado do Rio de Janeiro: Pieridae (Lepidoptera). Arquivos do Museu Nacional do Rio de Janeiro, 67 (3-4): 283-289.

Ribeiro, D. B. & A. V. L. Freitas. 2010. Differences in thermal responses in a fragmented landscape: Temperature affects the sampling of diurnal, but not nocturnal fruit-feeding Lepidoptera. The Journal of Research on the Lepidoptera 42: 1-4.

Ribeiro, D. B., P. I. Prado, K. S. Brown Jr. & A. V. L. Freitas. 2010. Temporal diversity patterns and phenology in fruit-feeding butterflies in the Atlantic Forest. Biotropica, 42(6): 710-716.

Freitas, A. V. L., R. B. Francini & T. S. Souza. 2009. Immature stages and natural history of the threatened butterfly Actinote quadra (Nymphalidae: Heliconiinae: Acraeini). Tropical Lepidoptera Research 19(2): 82-88.

Tourinho, J. L. & A. V. L. Freitas. 2009. Population biology of Euptoieta hegesia (Nymphalidae: Heliconiinae: Argynnini) in an urban area in Southeastern Brazil. The Journal of Research on the Lepidoptera, 41: 40-44.

Uehara-Prado. M. & A. V. L. Freitas. 2009. The effect of rainforest fragmentation on species diversity and mimicry ring composition of ithomiine butterflies. Insect Conservation and Diversity, 2: 23-28.

Uehara-Prado, M., J. O. Fernandes, A. M. Bello, G. Machado, A. J. Santos, F. Z. Vaz-de-Mello & A. V. L. Freitas. 2009. Selecting terrestrial arthropods as indicators of small-scale disturbance: a first approach in the Brazilian Atlantic Forest. Biological Conservation 142: 1220-1228.

Canuto, J. Z., T. A. Nogueira, K. S. Brown Jr. & A. V. L. Freitas. 2008. Lepidoptera, Nymphalidae, Ithomiinae, Scada reckia reckia: suggestion of recent range expansion throughout the State of Espirito Santo, SE Brazil. Checklist, 3(2): 159-160.

Ribeiro, D. B., P. I. Prado, K. S. Brown Jr. & A. V. L. Freitas. 2008. Additive partitioning of butterfly diversity in a fragmented landscape: Importance of scale and implications for conservation. Diversity and Distributions, 14: 961-968.

Azevedo, C. O., F. Z. Vaz-de-Mello, M. T. Tavares, A. D. Brescovit, P. de Marco Jr., K. Furieri, K. S. Brown Jr. & A. V. L. Freitas. 2007. Os invertebrados terrestres ameaçados de extinção no estado do Espírito Santo. Páginas 105-119 In: Espécies da fauna ameaçados de extinção no estado do Espírito Santo. Pasamani, M. & S. L. Mendes (organizadores.). IPEMA, Vitória, 140pp. (BOOK CHAPTER)

Uehara-Prado, M. K. S. Brown Jr. & A. V. L. Freitas. 2007. Species richness, composition and abundance of fruit-feeding butterflies in the Brazilian Atlantic Forest: comparison between a fragmented and a continuous landscape. Global Ecology and Biogeography, 16: 43-54.

Freitas, A. V. L. & C. Peña. 2006. Description of Genus Guaianaza for “Euptychiapronophila (Lepidoptera: Nymphalidae: Satyrinae) with a description of the immature stages. Zootaxa, 1163: 49-59.

Andrade, R. B. & A. V. L. Freitas. 2005. Population biology of two species of Heliconius (Nymphalidae: Heliconiinae) in a semi-deciduous forest in Southeastern Brazil. Journal of the Lepidopterists’ Society 59(4): 223-228.

Francini, R. B., A. V. L. Freitas & K. S. Brown Jr. 2005. Rediscovery of Actinote zikani (D’Almeida) (Nymphalidae, Heliconiinae, Acraeini): Natural history, population biology and conservation of an endangered butterfly in SE Brazil. Journal of the Lepidopterists’ Society 59(3): 134-142.

Freitas, A. V. L. 2005. Borboletas. Borboletas da Mata Atlântica. Publicação privada (1000 cópias) – Tuim Parque. 12 pp.

Uehara-Prado, M., A. V. L. Freitas & K. S. Brown Jr. 2005. Biological traits of frugivorous butterflies in a fragmented and a continuous landscape in the South Brazilian Atlantic Forest. Journal of the Lepidopterists’ Society 59(2): 96-106.

Francini, R. B., A. V. L. Freitas & C. M. Penz. 2004. Two new species of Actinote (Lepidoptera, Nymphalidae) from Southeastern Brazil. Zootaxa, 719: 1-10.

Freitas, A. V. L. 2004. A new species of Yphthimoides (Nymphalidae, Satyrinae) from southeastern Brazil. Journal of the Lepidopterists’ Society, 58(1): 7-12.

Uehara-Prado, M., A. V. L. Freitas, R. B. Francini & K. S. Brown Jr. 2004. Guia das borboletas frugívoras da Reserva Estadual do Morro Grande e região de Caucaia do Alto, Cotia (SP). Biota Neotropica, 4(1): 1-25.

Brown Jr., K. S. & A. V. L. Freitas. 2003. Butterfly Communities of Urban Forest Fragments in Campinas, São Paulo, Brazil: Structure, Instability, Environmental Correlates, and Conservation. Journal of Insect Conservation, 6(4): 217-231.

Freitas, A. V. L. 2002. Immature stages of Eteona tisiphone (Nymphalidae: Satyrinae). Journal of the Lepidopterists’ Society, 56(4): 286-288.

Freitas, A. V. L., J. Vasconcellos-Neto, F. Vanini, J. R. Trigo & K. S. Brown Jr. 2001. Population studies of Aeria olena and Tithorea harmonia (Nymphalidae, Ithomiinae) in Southeastern Brazil. Journal of the Lepidopterists’ Society 55(4): 150-157.

Freitas, A. V. L. & R. R. Ramos. 2001. Population biology of Parides anchises nephalion (Papilionidae) in a coastal site in Southeast Brazil. Brazilian Journal of Biology, 61(4): 623-630.

Brown Jr., K. S. & A. V. L. Freitas. 2000. Diversidade de Lepidoptera em Santa Teresa, Espírito Santo. Boletim do Museu de Biologia Mello Leitão, Nova Série, 11/12: 71-116.

Brown Jr., K. S. & A. V. L. Freitas. 2000. Atlantic Forest Butterflies: Indicators for Landscape Conservation. Biotropica, 32 (4b): 934-956.


Amazônia

Mota, L. L., S. J. Boddington, K. S. Brown Jr., C. J. Callaghan, G. Carter, W. Carter, S. M. Dantas, D. R. Dolibaina, K. Garwood, R. C. Hoyer, R. K. Robbins, A. Soh, K. R. Willmott & A. V. L. Freitas. 2022. The butterflies of Cristalino Lodge, in the Brazilian southern Amazonia: An updated species list with a significant contribution from citizen science. Biota Neotropica 22(3): e20221367. [PDF] [Vídeo]

Mota, L. L., A. H. B. Rosa, L. R. Vasconcellos, K. R. Willmott & A. V. L. Freitas. 2022. A new subspecies of Mechanitis lysimnia from southern Amazonia (Nymphalidae: Danainae: Ithomiini). Tropical Lepidoptera Research 32(1): 47-51. [PDF]

Rosser, N., L. T. Shirai, K. K. Dasmahapatra, J. Mallet & A. V. L. Freitas. 2020. The Amazon river is a suture zone for a polyphyletic group of co‐mimetic heliconiine butterflies. Ecography 43: 1–11. [PDF] [Vídeo]

Massardo, D., N. W. VanKuren, S. Nallu, R. R. Ramos, P. G. Ribeiro, K. L. Silva-Brandão, M. M. Brandão, M. B. Lion, A. V. L. Freitas, M. Z. Cardoso & M. R. Kronforst. The roles of hybridization and habitat fragmentation in the evolution of Brazil’s enigmatic longwing butterflies, Heliconius nattereri and H. hermathena. BMC Biology 18(1): 1-21. [PDF]

Rosser, N., A. V. L. Freitas, B. Huertas, M. Joron, G. Lamas, C. Mérot, F. Simpson, K. R. Willmott, J. Mallet & K. K. Dasmahapatra. 2019. Cryptic speciation associated with geographic and ecological divergence in two Amazonian Heliconius butterflies. Zoological Journal of the Linnean Society 186(1): 233-249.

Freitas, A. V. L., R. R. Ramos, K. L. Silva-Brandão, N. Coutoné, L. M. Magaldi, J. L. Pablos, N. Rosser & K. S. Brown Jr. 2019. A New Subspecies of Heliconius hermathena (Nymphalidae: Heliconiinae) from Southern Amazonia. Neotropical Entomology 48(3): 467-475.

Freitas, A. V. L., L. L. Mota, E. P. Barbosa & J. Y. O. Carreira. 2019. Immature stages of the Selaginella-feeding Euptychia mollina (Nymphalidae: Satyrinae) from the Brazilian Amazon. Zoologia, 36: e32053. [PDF]

Freitas, A. V. L., L. L. Mota, T. Zacca & E. P. Barbosa. 2019. Description of a new and highly distinctive genus and species of Euptychiina (Lepidoptera: Nymphalidae: Satyrinae) from the Brazilian southern Amazon. Revista Brasileira de Entomologia 63(3): 254-261.

de Andrade, R. B., J. K. Balch, J. Y. O. Carreira, P. M. Brando & A. V. L. Freitas. 2017. The impacts of recurrent fires on diversity of fruit-feeding butterflies in a south-eastern Amazon forest. Journal of Tropical Ecology 33 (1): 22-32. [PDF]

Seixas, R. R., S. E. Santos, Y. Okada & A. V. L. Freitas. 2017. Population biology of the sand forest specialist butterfly Heliconius hermathena hermathena (Hewitson) (Nymphalidae: Heliconiinae) in central Amazonia. Journal of the Lepidopterists’ Society 71(3): 133–140.

Ribeiro, D. B., M. R. Williams, A. Specht & A. V. L. Freitas. 2016. Vertical and temporal variability in the probability of detection of fruit-feeding butterflies and moths (Lepidoptera) in tropical forest. Austral Entomology 55(1): 112-120.

Santos, J. P., A. V. L. Freitas, P. A. L. Constantino & M. Uehara-Prado. 2014. Guia de identificação de tribos de borboletas frugívoras. Amazônia. In. Pereira, A. B. & P. A. L. Constantino (Eds.). Monitoramento de Biodiversidade. MMA/ICMBio/GIZ. 12 pp.

Ribeiro, D. B. & A. V. L. Freitas. 2012. The effect of reduced-impact logging on fruit-feeding butterflies in Central Amazon, Brazil. Journal of Insect Conservation, 16: 733-744.

Raimundo, R. L. G., A. V. L. Freitas, R. N. S. Costa, J. B. F. Oliveira, A. B. Melo & K. S. Brown Jr. 2003. Manual de monitoramento ambiental usando borboletas e libélulas – Reserva Extrativista do Alto Juruá, 36 pp. (il). Série Pesquisa e Monitoramento Participativo em Áreas de Conservação Gerenciadas por Populações Tradicionais, Volume 1. Campinas, CERES/Laboratório de Antropologia e Ambiente. ISBN: 85-86572-13-6 (BOOK)

Brown Jr., K. S. & A. V. L. Freitas. 2002. Diversidade Biológica no Alto Juruá: Avaliação, Causas e Manutenção. Páginas 33-42; pranchas 13-16 In: Enciclopédia da floresta. O Alto Juruá: Práticas e conhecimentos das populações. Carneiro da Cunha, M. M. & M. B. Almeida (organizadores). São Paulo: Companhia das Letras, 735pp. (BOOK CHAPTER)


Cerrado

Silva, L. D., A. H. B. Rosa, A. V. L. Freitas, C. A. Iserhard, L. A. Kaminski, O. J. Marini-Filho, B. M. Thompson, G. R. Soares & M. M. de Sousa. 2022. Papilionoidea inventory of the Sempre Vivas National Park, Minas Gerais State, southeastern Brazil (Insecta: Lepidoptera). SHILAP Revista de Lepidopterología 50: 367-380. [PDF]

Freitas, A. V. L., D. G. Muniz, J. Y. O. Carreira & P. S. Oliveira. 2021. Fruit-Feeding Butterfly Assemblages in a Neotropical Savanna: Assessing Phenological Patterns Using Baited Traps. The Journal of the Lepidopterists’ Society 75(2): 88-103. [PDF]

Freire, G. B., T. Silva, H. Oliveira, C. Collier, H. P. Rodrigues, J. P. Dias, J. P. Santos, O. J. Marini-Filho, A. V. L. Freitas, A. M. Smilanich, L. A. Dyer & Ivone R. Diniz. 2021. Good Things Come in Larger Packages: Size Matters for Adult Fruit-Feeding Butterfly Dispersal and Larval Diet Breadth. Diversity 13(12): 664. [PDF]

Nascimento, A.R., I. K. Malinov, G. Freire Jr, A. V. L. Freitas & I. R. Diniz. 2020. The Temporal Dynamics of Two Morpho Fabricius, 1807 Species (Lepidoptera: Nymphalidae) are Affected Differently by Fire in the Brazilian Savanna. Environmental Entomology 49(6): 1449–1454. [PDF]

Rosa, A. H. B., L. V. Perillo, F. S. Neves, D. B. Ribeiro & A. V. L. Freitas. 2019. Butterflies collected using malaise traps as useful bycatches for ecology and conservation. Journal of Threatened Taxa 11(9): 14235-14237. [PDF]

Freitas, A. V. L., P. E. Gueratto & A. R. Nascimento. 2019. New records of high altitude Actinote (Nymphalidae: Heliconiinae) from central Brazil extend the known distribution by 650 km. Journal of the Lepidopterists’ Society 73(3): 208-210.

Freitas A.V.L., Barbosa E.P., Zacca T., Marín M.A., Beirão M.V., Silva A.R.M., Casagrande M., Espeland M. & Willmott K.R. 2018. Before it is too late: description of a new genus and species of butterfly from a highly threatened Brazilian biome. Revista Brasileira de Entomologia 62: 148–158. [PDF]

Kaminski, L. A., C. J. Callaghan, N. Seraphim, L. M. Magaldi, L. Volkmann & A. V. L. Freitas. 2017. Sertania gen. nov., a new genus of butterflies (Lepidoptera: Riodinidae) from the South American dry diagonal. Zootaxa 4312 (1): 165-179.

Santos, J. P., A. V. L. Freitas, P. A. L. Constantino & M. Uehara-Prado. 2014. Guia de identificação de tribos de borboletas frugívoras. Cerrado. In. Pereira, A. B. & P. A. L. Constantino (Eds.). Monitoramento de Biodiversidade. MMA/ICMBio/GIZ. 12 pp.

Muniz, D. G., A. V. L. Freitas & P. S. Oliveira. 2012. Phenological relationships of Eunica bechina (Lepidoptera: Nymphalidae) and its host plant, Caryocar brasiliense (Caryocaraceae), in a Neotropical savanna. Studies in Neotropical Fauna and Environment 47(2): 111-118.

Freitas, A. V. L., E. O. Emery & O. H. H. Mielke. 2010. A new species of Moneuptychia Forster (Lepidoptera: Satyrinae: Euptychiina) from Central Brazil. Neotropical Entomology 39(1): 83-90.

Kaminski, L. A., A. V. L. Freitas & P. S. Oliveira. 2010. Interaction between mutualisms: Ant-tended butterflies exploit enemy-free space provided by ant-treehopper associations. The American Naturalist, 176(3): 322-334.

Rodrigues, D., L. A. Kaminski, A. V. L. Freitas & P. S. Oliveira. 2010. Trade-offs underlying polyphagy in a facultative ant-tended florivorous butterfly: the role of host plant quality and enemy-free space. Oecologia, 163: 719-728.

Kaminski, L. A., S. F. Sendoya, A. V. L. Freitas & P. S. Oliveira. 2009. Ecologia comportamental na interface formiga-planta-herbívoro: interações entre formigas e lepidópteros. Oecologia Brasiliensis, 13(1): 27-44.

Sendoya, S. F., A. V. L. Freitas & P. S. Oliveira. 2009. Egg-laying butterflies distinguish predaceous ants by sight. The American Naturalist 174(1): 134-140.

Oliveira, P. S. & A. V. L. Freitas. 2004. Ant-Plant-Herbivore Interactions in the Neotropical Cerrado Savanna. Review. Naturwissenschaften, 91(12): 557-570.

Oliveira, P. S., A. V. L. Freitas & K. Del-Claro. 2002. Ant foraging on plant foliage: contrasting effects on the behavioral ecology of insect herbivores. Páginas 287-305 In: The Cerrados of Brazil: Ecology and Natural History of a Neotropical Savanna. Oliveira, P. S. & R. J. Marquis (editores). Columbia University Press, NY, viii + 398pp. (BOOK CHAPTER)


Caatinga

Melo, D. H. A., A. V. L. Freitas, M. Tabarelli, B. K. C. Filgueiras & I. R. Leal. 2022. Aridity and chronic anthropogenic disturbance as organizing forces of fruit-feeding butterfly assemblages in a Caatinga dry forest. Biotropica, 00, 1–12. [PDF]


Cordilheira dos Andes

Nakahara, S., M. Rodríguez-Melgarejo, K. Kleckner, T. Corahua-Espinoza, R. Tejeira, M. Espeland, M. M. Casagrande, E. P. Barbosa, J. See, G. Gallice, G. Lamas & K. R. Willmott. 2022. Systematic Revision of a New Butterfly Genus, Cisandina Nakahara & Espeland, n. gen., with Descriptions of Three New Taxa (Lepidoptera: Nymphalidae: Satyrinae). Insect Systematics and Diversity 6(1): 2; 1–30. [PDF]

Álvares, C. F., A. Clavijo-Giraldo, S. I. Uribe, T. W. Pyrcz, C. A. Iserhard, A. V. L. Freitas & M. A. Marín. 2022. Sampling performance of bait traps in high Andean fruit-feeding butterflies. Neotropical Biodiversity 7(1): 507-513. [PDF]

Marín, M. A., A. López-Rubio, A. Clavijo, T. W. Pyrcz, A. V. L. Freitas, S. I. Uribe & C. F. Álvarez. 2021. Use of species delimitation approaches to tackle the cryptic diversity of an assemblage of high Andean butterflies (Lepidoptera: Papilionoidea). Genome. [PDF]

Nakahara, S., E. P. Barbosa, I. Nakamura, G. Lamas & A. V. L. Freitas. 2021. Description of a new species of Yphthimoides Forster, 1964 from Peru (Lepidoptera: Nymphalidae: Satyrinae). Tropical Lepidoptera Research 31(3):138-144. [PDF]

Willmott, K. R., M. A. Marín, S. Nakahara, T. Pomerantz, G. Lamas, B. Huertas, M. Espeland, L. Xiao., J. P. W. Hall, J. I. R. Willmott & A. V. L. Freitas. 2019. A revision of the new Andean butterfly genus Optimandes Marín, Nakahara & Willmott, n. gen., with the description of a new species (Nymphalidae: Satyrinae: Euptychiina). Tropical Lepidoptera Research 29(1): 29-44.

Willmott, K. R., A. V. L. Freitas, J. P. W. Hall, K. L. Silva-Brandão & M. Paluch. 2009. A new species of Actinote Hübner from the Eastern Andes of Ecuador (Lepidoptera: Nymphalidae: Heliconiinae). Proceedings of the Entomological Society of Washington, 111(1): 47-56.

Algarin, C. M., M. A. Marin, S. Uribe & A. V. L. Freitas. 2008. Tabla de vida de Pseudodirphia pallida (Lepidoptera: Saturniidae) en condiciones de laboratório. Revista Colombiana de Entomologia, 34(1): 116-120.


Divulgação científica

Freitas, A. V. L., Zacca, T.; Moraes, S. S.; Aguiar, T. M. C.; Barbosa, E. P. 2018. Muito além de um simples acervo. Coletiva (Revista eletrônica de divulgação científica) – Diversidade Socioambiental, v. 3., 21 de setembro de 2018. [PDF]

Rodrigues, L. & A. V. L. Freitas. 2004. Borboletas: delicada beleza. Obra em 4 volumes. Lemos Editorial & Gráficos Ltda/Biosintética, São Paulo. 184pp. ISBN: 85-7450-170-0 (BOOK)


Ensino de Biologia

D´Ambrosio, M., A. V. L. Freitas & F. S. Santos. 2017. Investigating how undergraduate Biological Sciences students understand tree-thinking: results from two Brazilian institutions. Conexão Ciência 12(número especial 2): 204-209.